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M
Mucilaginous
Mucilaginous.
Resembling mucilage: moist, ropy, slimy and sticky.
TWS
Mucro
Mucro.
Minute
awn
or
excurrent
mid-
nerve
of an organ.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
minute
Minute.
A small measure of time.
TWS
Minute.
A small thing.
TWS
awn, awns
Awn, arista.
Slender, straight or bent bristle. In grasses: usually a continuation of the mid-nerve of lemmas or glumes.
2020 International Rules for Seed Testing (ISTA)
Awn.
The slender bristle extending from the tip or back of the lemma or the glume of a grass spikelet.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil anad Crop Sciences, Colorado State University. Pages 104-11
excurrent
Excurrent.
Running out as a nerve of a leaf running out beyond the margin.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
nerve, nerves
Nerve.
Vascular veins of the blades, glumes and lemmas.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Multiple fruit
Multiple fruit.
A
fruit
which develops from a complete
inflorescence
, as opposed to a true or
simple fruit
, which develops from one flower.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
fruit
Fruit.
The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Fruit.
In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity.
SCST Seed Technologist Training Manual, 2018
Fruit.
The ripened ovary of a seed plant and associated parts.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
inflorescence
Inflorescence.
A flower cluster.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Inflorescence.
The portion of the plant adapted for flowering and fruiting.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
simple fruit
Simple fruit.
A fruit which develops from the gynoecium of a single flower (=true fruit).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Multiple seed unit
Multiple seed unit.
Spikelet
or part of spikelet with more than one
floret
, with or without
glumes
.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Multiple unit.
Is a
seed unit
that includes one or more structures as follows: (1) An attached
sterile
or
fertile floret
that extends to or beyond the tip of a
fertile
floret; (2) A fertile floret with basally attached
glume
, glumes, or basally attached
sterile floret
of any length; (3) A fertile floret with two or more attached sterile and/or
fertile florets
of any length.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
spikelet, spikelets
Spikelet.
The unit of a grass inflorescence comprising one or more florets subtended by one or two sterile glumes. For the purposes of the Rules, the term spikelet includes, as well as a fertile floret, either one or more additional fertile or completely infertile florets, or glumes.
2020
International Rules for Seed Testing (ISTA)
Spikelet.
Part of a grass inflorescence including one or more florets subtended by one or two glumes.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Spikelet.
In grasses: One or more attached florets usually subtended by one or more bracts (glumes).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Spikelet.
The unit of a grass inflorescence that consists of a pair of empty glumes that enclose one of more florets arranged around a rachilla.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
floret, florets
Floret.
General: An individual flower within a cluster. In grasses: A flower usually enclosed by two bracts (lemma and palea).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Floret.
A flower within an inflorescence or in a grass spikelet.
SCST Seed Technologist Training Manual, 2018
Floret.
The lemma and palea with enclosed pistil and stamens, or, in
Poaceae
, the mature caryopsis; for the purpose of the Rules, the term floret refers to the fertile floret, with or without additional sterile lemmas.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Floret.
The lemma and palea with enclosed pistil and stamens or the mature caryopsis in Poaceae; for the purpose of the Rules, the term floret refers to the fertile floret with or without additional sterile lemmas.
2020
International Rules for Seed Testing (ISTA)
Floret.
In grasses, a flower consisting of lemma and palea which enclose the flowering parts -- stamens, pistil and lodicules. May be perfect, staminate, pistillate, neuter, sterile, and so on. A small flower in other plant families.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
glume, glumes
Glume.
In grasses: A bract—often paired—at the base of a spikelet.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Glume.
One of the two usually sterile bracts at the base of a grass spikelet.
2020
International Rules for Seed Testing (ISTA)
Glumes.
The pair of empty bracts at the base of a spikelet.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
seed unit
Seed unit.
Commonly found dispersal unit, i.e. achenes and similar fruits, schizocarps, florets etc., as defined for each genus or species in the ISTA Rules, Table 3B Parts 1 & 2.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed unit.
The structure usually regarded as a seed in planting practices and in commercial channels. Refer to section 3.2 e for pure seed unit definitions.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Seed Unit.
Commonly found dispersal unit, i.e. achenes and similar fruits, schizocarps, florets etc, as defined for each genus or species in the Pure Seed Definitions in Table 3B parts 1 and 2.
2020 International Rules for Seed Testing (ISTA)
sterile
Sterile.
Without functional sex organs (for grass florets: without caryopsis).
2020
International Rules for Seed Testing (ISTA)
Sterile.
Without pistils. A sterile floret may be staminate or neuter. It may even lack a palea, and consist of nothing but a lemma.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
fertile floret, fertile florets
Fertile floret.
A floret that encloses a caryopsis.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
fertile
Fertile.
With functional sex organs; (for grass florets: having a caryopsis).
2020
International Rules for Seed Testing (ISTA)
Fertile.
With functional sex organs.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Fertile.
Capable of producing fruit; having pistils. A fertile floret may be pistillate or perfect.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
glume, glumes
Glume.
In grasses: A bract—often paired—at the base of a spikelet.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Glume.
One of the two usually sterile bracts at the base of a grass spikelet.
2020
International Rules for Seed Testing (ISTA)
Glumes.
The pair of empty bracts at the base of a spikelet.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
sterile floret, sterile florets
Sterile floret.
A floret with no caryopsis.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
fertile floret, fertile florets
Fertile floret.
A floret that encloses a caryopsis.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Multiple unit procedure
Multiple unit procedure.
A purity procedure used to determine the amount of
inert
material in
spikelets
and
florets
that do not
disarticulate
in certain prescribed grasses by means of a mathematical factor method.
SCST Seed Technologist Training Manual, 2018
inert matter, inert
Inert matter.
Inert matter shall include seeds and seed-like structures from both crop and weed plants and other materials not described in section 3.2 and Table 3A or can be described as follows: seeds and seed-like structure from crop and weed plants and other non-seed matter.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
spikelet, spikelets
Spikelet.
The unit of a grass inflorescence comprising one or more florets subtended by one or two sterile glumes. For the purposes of the Rules, the term spikelet includes, as well as a fertile floret, either one or more additional fertile or completely infertile florets, or glumes.
2020
International Rules for Seed Testing (ISTA)
Spikelet.
Part of a grass inflorescence including one or more florets subtended by one or two glumes.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Spikelet.
In grasses: One or more attached florets usually subtended by one or more bracts (glumes).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Spikelet.
The unit of a grass inflorescence that consists of a pair of empty glumes that enclose one of more florets arranged around a rachilla.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
floret, florets
Floret.
General: An individual flower within a cluster. In grasses: A flower usually enclosed by two bracts (lemma and palea).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Floret.
A flower within an inflorescence or in a grass spikelet.
SCST Seed Technologist Training Manual, 2018
Floret.
The lemma and palea with enclosed pistil and stamens, or, in
Poaceae
, the mature caryopsis; for the purpose of the Rules, the term floret refers to the fertile floret, with or without additional sterile lemmas.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Floret.
The lemma and palea with enclosed pistil and stamens or the mature caryopsis in Poaceae; for the purpose of the Rules, the term floret refers to the fertile floret with or without additional sterile lemmas.
2020
International Rules for Seed Testing (ISTA)
Floret.
In grasses, a flower consisting of lemma and palea which enclose the flowering parts -- stamens, pistil and lodicules. May be perfect, staminate, pistillate, neuter, sterile, and so on. A small flower in other plant families.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
disarticulate
Disarticulate.
Separate at a joint at maturity (see articulate).
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
N
Navicular
Navicular.
Boat-shaped.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Necrosis
Necrosis.
Dead or deteriorating
seedling
tissue, that may be caused by injury, disease or physiological breakdown.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
seedling, seedlings
Seedling.
A young plant developing from the embryo of a seed.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Negative geotropism
Negative geotropism.
Negative geotropism
is caused by a physiological disorder usually characterized by
root
structures that grow upward.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
negative geotropism
Negative geotropism.
Negative geotropism is caused by a physiological disorder usually characterized by root structures that grow upward.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
root, roots
Root.
In dicotyledons and gymnosperms, the root system serves three major functions: (1) to anchor the plant in the soil, (2) to absorb water and dissolved salts from the soil and (3) to conduct the water and salts to the hypocotyl, cotyledons and epicotyl. The embryonic root, or radicle, is located at the basal end of the embryo and is usually the first seedling structure to rupture the testa. After emergence it is referred to as the primary root. The primary root elongates rapidly and soon numerous root hairs develop, greatly increasing the absorbing surface of the roots. As the seedling continues to grow, secondary roots develop from the primary root and from other secondary roots. Roots may also emerge from other structures (e.g. the hypocotyl) and are referred to as adventitious roots.
As in the dicotyledons, the monocotyledon root system serves to anchor the plant in soil, absorb water and dissolved salts from the soil and to conduct the water and salts to the growing seedling. The embryonic root, or radicle, is situated at the basal end of the embryo and, in the case of the
Poaceae
, its apex is covered by the coleorhiza. After the radicle emerges it is referred to as the primary root. In some species of the
Poaceae
(e.g.
Triticum
) the primary root is indistinguishable from the other roots that develop from the scutellar node region and hence all of these are referred to as seminal roots. Roots that develop from structures above the scutellar or cotyledonary node are called adventitious roots. Secondary roots may develop from seminal and adventitious roots.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Root.
A portion of a higher plant bearing neither leaves nor reproductive organs, usually underground.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Nerve, nerves
Nerve.
Vascular
veins
of the blades,
glumes
and
lemmas
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
vein, veins
Vein.
A bundle of threads of fibro-vascular tissue in a leaf or other organ.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Veins.
Strands of vascular tissue visible from the surface of plant structures.
SCST Seed Technologist Training Manual, 2018
glume, glumes
Glume.
In grasses: A bract—often paired—at the base of a spikelet.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Glume.
One of the two usually sterile bracts at the base of a grass spikelet.
2020
International Rules for Seed Testing (ISTA)
Glumes.
The pair of empty bracts at the base of a spikelet.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
lemma, lemmas
Lemma.
The outer (lower) bract of a grass floret, sometimes referred to as the flowering glume or the lower or outer palea. Bract enclosing the caryopsis on the outer (dorsal) side.
2020
International Rules for Seed Testing (ISTA)
Lemma.
The outer (lower) bract of a grass floret; enclosing the caryopsis on the outer (dorcal) side.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Lemma.
The lower of two bracts that subtend a grass flower.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Lemma.
The lower of the two bracts enclosing the flower in the spikelet of grasses.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Nerved
Nerved.
Having pronounced
veins
.
SCST Seed Technologist Training Manual, 2018
vein, veins
Vein.
A bundle of threads of fibro-vascular tissue in a leaf or other organ.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Veins.
Strands of vascular tissue visible from the surface of plant structures.
SCST Seed Technologist Training Manual, 2018
Neuter
Neuter.
Without stamens or pistils.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Nodding
Nodding.
Inclined from the vertical.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Node, nodes
Node.
The point on a stem from which a
leaf
arises.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
leaf, leaves
Leaf.
Lateral organ of the stem.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Nodule, nodules
Nodule.
An enlargement on the
roots
of
legumes
within which the nitrogen-fixing bacteria live.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
root, roots
Root.
In dicotyledons and gymnosperms, the root system serves three major functions: (1) to anchor the plant in the soil, (2) to absorb water and dissolved salts from the soil and (3) to conduct the water and salts to the hypocotyl, cotyledons and epicotyl. The embryonic root, or radicle, is located at the basal end of the embryo and is usually the first seedling structure to rupture the testa. After emergence it is referred to as the primary root. The primary root elongates rapidly and soon numerous root hairs develop, greatly increasing the absorbing surface of the roots. As the seedling continues to grow, secondary roots develop from the primary root and from other secondary roots. Roots may also emerge from other structures (e.g. the hypocotyl) and are referred to as adventitious roots.
As in the dicotyledons, the monocotyledon root system serves to anchor the plant in soil, absorb water and dissolved salts from the soil and to conduct the water and salts to the growing seedling. The embryonic root, or radicle, is situated at the basal end of the embryo and, in the case of the
Poaceae
, its apex is covered by the coleorhiza. After the radicle emerges it is referred to as the primary root. In some species of the
Poaceae
(e.g.
Triticum
) the primary root is indistinguishable from the other roots that develop from the scutellar node region and hence all of these are referred to as seminal roots. Roots that develop from structures above the scutellar or cotyledonary node are called adventitious roots. Secondary roots may develop from seminal and adventitious roots.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Root.
A portion of a higher plant bearing neither leaves nor reproductive organs, usually underground.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
legume, legumes
Legume.
A dry fruit consisting of one carpel, splitting by two longitudinal sutures with a row of seeds on the inner side of the central suture; pod, as in
Fabaceae
.
SCST Seed Technologist Training Manual, 2018
Normal seedling, normal, normals
Normal seedling.
A
seedling
with all
essential structures
present and capable of developing into a plant under favorable conditions; certain defects may be present if they are judged to be not so severe as to impede further development of the plant (see
abnormal seedling
).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
seedling, seedlings
Seedling.
A young plant developing from the embryo of a seed.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
essential structure, essential structures
Essential structure.
Structure which is critical for continued development of the seedling into a plant.
SCST Seed Technologist Training Manual. 2018
Essential structure.
Any seedling structure that must be considered when classifying a seedling as either normal or abnormal. In general, an essential structure is one that is critical for successful establishment and development of a seedling into a plant. Structure which is critical for continued development of the seedling into a plant.
Baalbaki, R.Z. Germination & Dormancy. TWS Website. 2020
abnormal seedling, abnormals
Abnormal seedling.
A seedling that does not have all the essential structures or is damaged, deformed or decayed to such an extent that normal development is prevented (see normal seedling).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Noxious working sample
Noxious weed seed examination working sample.
The sub-sample taken from the submitted sample on which the noxious
weed seed
examination is performed. Refer to sections 2.3 b and 5.1.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
weed seed
Weed seed.
Seeds, florets, bulblets, tubers, or sporocarps of plants recognized as weeds by laws, official regulations, or by general usage shall be considered weed seeds; refer to section 4.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Weeds.
Undesirable species that are excessively competitive, difficult to control or eradicate, poisonous, or simply not wanted.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 3, 2019
Nucellus
Nucellus.
The
ovule
tissue within the
integuments
around the
embryo sac
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
ovule
Ovule.
The immature seed within the ovary.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Ovule.
A structure, consisting of a female gametophyte, nucellus, and integuments, which develops into a seed after fertilization.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
integument, integuments
Integument.
The envelope of an ovule which becomes the seed coat or testa (generally two integuments present). In coniferous seeds integument also refers to the tissue attaching the wing to the seed.
2020
International Rules for Seed Testing (ISTA)
Integument.
The envelope of an ovule, which becomes the testa.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Integuments.
Outermost coverings of an ovule.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
embryo sac
Embryo sac.
A large thin-walled cell within the ovule in which the embryo develops after fertilization.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Nut, nuts
Nut.
A dry
indehiscent
fruit
with a hard wall that is usually shed as a one-seeded unit. It forms from more than one
carpel
, but only one
seed
develops, the rest aborting. The
pericarp
is usually lignified and is often partially or completely surrounded by a cupule. True
nuts
include the acorn
(Quercus)
, hazelnut
(Corylus)
and beechnut
(Fagus).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Nut.
A hard, dry, indehiscent fruit, usually single-seeded.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
indehiscent
Indehiscent.
Not opening; fruits which do not open at maturity.
2020
International Rules for Seed Testing (ISTA)
Indehiscent.
Fruits in which the fruit wall does not split open at maturity to release the seed (see dehiscent).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Indehiscent fruit.
A fruit that does not open at maturity.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Indehiscent.
Fruits remaining closed at maturity.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
fruit
Fruit.
The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Fruit.
In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity.
SCST Seed Technologist Training Manual, 2018
Fruit.
The ripened ovary of a seed plant and associated parts.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
carpel, carpels
Carpel.
The female reproductive organ of flowering plants. It consists of an ovary, containing one or more ovules (which became seeds after fertilization), and a stigma, a surface receptive for pollen grains.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Carpel.
A simple pistil or an element of a compound pistil.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
pericarp, pericarps
Pericarp
[
fruit coat
]. The wall of the mature ovary or fruit.
2020
International Rules for Seed Testing (ISTA)
Pericarp.
The ovary wall. the fruit coat enclosing the seed.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Pericarp.
Fruit wall.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Pericarp.
The wall of a ripened ovary; the fruit coat.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
nut, nuts
Nut.
A dry indehiscent fruit with a hard wall that is usually shed as a one-seeded unit. It forms from more than one carpel, but only one seed develops, the rest aborting. The pericarp is usually lignified and is often partially or completely surrounded by a cupule. True nuts include the acorn
(Quercus)
, hazelnut
(Corylus)
and beechnut
(Fagus).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Nut.
A hard, dry, indehiscent fruit, usually single-seeded.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Nutlet, nutlets
Nutlet.
A small
nut
.
2020
International Rules for Seed Testing (ISTA)
Nutlet.
A small nut, a one seeded portion of a
fruit
originating from half of a
carpel
(e.g.
Boraginaceae
,
Lamiaceae
and
Verbinaceae
).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Nutlet.
A dry, one-seeded,
indehiscent
section of a fruit in the
Boraginaceae
,
Lamiaceae
and
Verbenaceae
.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
nut, nuts
Nut.
A dry indehiscent fruit with a hard wall that is usually shed as a one-seeded unit. It forms from more than one carpel, but only one seed develops, the rest aborting. The pericarp is usually lignified and is often partially or completely surrounded by a cupule. True nuts include the acorn
(Quercus)
, hazelnut
(Corylus)
and beechnut
(Fagus).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Nut.
A hard, dry, indehiscent fruit, usually single-seeded.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
fruit
Fruit.
The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Fruit.
In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity.
SCST Seed Technologist Training Manual, 2018
Fruit.
The ripened ovary of a seed plant and associated parts.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
carpel, carpels
Carpel.
The female reproductive organ of flowering plants. It consists of an ovary, containing one or more ovules (which became seeds after fertilization), and a stigma, a surface receptive for pollen grains.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Carpel.
A simple pistil or an element of a compound pistil.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
indehiscent
Indehiscent.
Not opening; fruits which do not open at maturity.
2020
International Rules for Seed Testing (ISTA)
Indehiscent.
Fruits in which the fruit wall does not split open at maturity to release the seed (see dehiscent).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Indehiscent fruit.
A fruit that does not open at maturity.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Indehiscent.
Fruits remaining closed at maturity.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
O
Obcordate
Obcordate.
Heart-shaped with broad end at the tip.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Oblanceolate
Oblanceolate.
Lanceolate
with broadest part at the
apex
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
lanceolate
Lanceolate.
Several times longer than wide, tapering from base to apex; lance-shaped.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
apex
Apex.
The tip, point, or angular summit of a structure.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Oblique
Oblique.
Slanting or inclined.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Oblong
Oblong.
Longer than broad with sides nearly parallel.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Obovate
Obovate.
Egg-shaped outline with broadest end above.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Obsolete
Obsolete.
Greatly reduced in size so as to be difficult to see.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Obsolete.
Almost absent or
vestigial
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
vestigial
Vestigial.
A small, imperfectly developed part.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Obtuse
Obtuse.
Blunt or rounded.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Open
Open.
Loose;
opposite
of
dense
or compact.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
opposite
Opposite.
Bearing two leaves or buds at a node on opposite sides of a stem.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
dense
Dense.
Parts massed or crowded together.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Opposite
Opposite.
Bearing two
leaves
or
buds
at a
node
on
opposite
sides of a stem.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
leaf, leaves
Leaf.
Lateral organ of the stem.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
bud, buds
Bud.
An unexpanded flower or a rudimentary leaf, stem, or branch.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
node, nodes
Node.
The point on a stem from which a leaf arises.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
opposite
Opposite.
Bearing two leaves or buds at a node on opposite sides of a stem.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Order
Order.
A category in the classification of plants ranking below class and above family; an
order
is composed of families.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
order
Order.
A category in the classification of plants ranking below class and above family; an order is composed of families.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Orthodox
Orthodox.
Desiccation tolerant. Capable of surviving drying to very low moisture contents.
Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020
Other crop seed
Other crop seed.
Seeds
of plants grown as crops (other than the kind(s) and cultivar(s) included in the pure
seed
) shall be considered other crop seeds, unless recognized as weed seeds by laws, regulations, or by general usage; refer to section 4.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Other crop.
Species
that are usually involved in seed commerce but are not intended to be part of the
seed lot
being tested. Contamination by these seeds is undesirable in the seed lot but not usually harmful.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 3, 2019
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
species
Species.
A category of classification lower than a genus that is made up of plants which possess in common distinctive characteristics that are reproduced in their offspring. The species name included second to the genus in the scientific binomial.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
seed lot
Seed lot.
A seed lot is a specified quantity of seed that is physically and uniquely identified.
2020
International Rules for Seed Testing, (ISTA)
Lot of seed.
The term "lot of seed" means a definite quantity of seed identified by a number, every portion or bag which is uniform, within permitted tolerances, for the factors which appear in the labeling.
Agricultural Marketing Service, USDA. Federal Seed Act, Part 201. Federal Seed Act Regulations. 201.2 Terms Defined. Current as of May 21, 2020.
Other seeds
Other seeds.
Other seeds
shall include
seed
units of any plant
species
other than that of pure seed.
2020
International Rules for Seed Testing, (ISTA)
other seeds
Other seeds.
Other seeds shall include seed units of any plant species other than that of pure seed.
2020
International Rules for Seed Testing, (ISTA)
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
species
Species.
A category of classification lower than a genus that is made up of plants which possess in common distinctive characteristics that are reproduced in their offspring. The species name included second to the genus in the scientific binomial.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Ovary
Ovary.
The
basal
, expanded part of the
pistil
that contains the ovules.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Ovary.
The basal enlarged portion of a pistil within which
seeds
develop.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
basal
Basal.
At the base or bottom.
SCST Seed Technologist Training Manual, 2018
pistil
Pistil.
A part (or whole) of the gynoecium, consisting of either a separate, free carpel or two or more fused carpels. A typical pistil comprises an ovary, a style and a stigma.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Pistil.
The ovule-bearing portion of a flower consisting of a stigma, style and ovary.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Ovate
Ovate.
Egg-shaped, broadest part at base.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Oven test method
Oven test method.
Seed
moisture test in which a quantity of seed is weighed prior to and after drying in an oven at a prescribed temperature for a specified time and the loss in weight is calculated as percentage moisture content on a fresh or dry weight basis.
SCST Seed Technologist Training Manual, 2018
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Ovule
Ovule.
The immature
seed
within the
ovary
.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Ovule.
A structure, consisting of a
female gametophyte
,
nucellus
, and
integuments
, which develops into a seed after
fertilization
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
ovary
Ovary.
The basal, expanded part of the pistil that contains the ovules.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Ovary.
The basal enlarged portion of a pistil within which seeds develop.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
female gametophyte
Female gametophyte.
In gymnosperms the nutritive tissue is the mature female gametophyte, sometimes also referred to as the primary endosperm because it is already present before fertilization.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
nucellus
Nucellus.
The ovule tissue within the integuments around the embryo sac.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
integument, integuments
Integument.
The envelope of an ovule which becomes the seed coat or testa (generally two integuments present). In coniferous seeds integument also refers to the tissue attaching the wing to the seed.
2020
International Rules for Seed Testing (ISTA)
Integument.
The envelope of an ovule, which becomes the testa.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Integuments.
Outermost coverings of an ovule.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
fertilization
Fertilization.
A sexual process in which two dissimilar gametes fuse.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Ovulode
Ovulode.
An
ovule
that did not develop into a mature
seed
, as in
Eucalyptus spp
.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
ovule
Ovule.
The immature seed within the ovary.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Ovule.
A structure, consisting of a female gametophyte, nucellus, and integuments, which develops into a seed after fertilization.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
P
Paired tests
Paired test.
Test procedures used on
seeds
having an unknown degree of
dormancy
. Samples are tested both with and without
prechill
or other treatments prescribed for breaking dormancy. Refer to section 6.9 n (3) for trees and shrubs.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
seed, seeds
Seed.
The part of a plant which is able to develop into a new plant.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Seed.
Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Seed.
The ripened ovule, enclosing the rudimentary plant and food necessary for its germination.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
dormant seeds, dormancy
Dormant seeds.
Viable seeds, other than hard seeds, which fail to germinate when provided the specified germination conditions for the kind of seed in question.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
Dormancy.
Delayed germination or growth; a condition of inactivity.
SCST Seed Technologist Training Manual, 2018
prechill
Prechill.
A cold, moist treatment applied to seeds to overcome dormancy prior to the germination test. The prechill method varies among species, but is usually performed by holding imbibed seeds at a low temperature for a specified period of time.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Palea
Palea.
The upper (inner)
bract
of a grass
floret
, sometimes called the inner or upper
palea
. Bract enclosing the
caryopsis
on the inner
ventral
side.
2020
International Rules for Seed Testing (ISTA)
Palea.
The upper (inner) bract of a grass floret, the bract enclosing the caryopsis on the inner (ventral) side.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Palea.
The uppermost of two bracts subtending a grass flower.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Palea.
The upper of the two bracts enclosing the flower in the
spikelet
of grasses.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
bract
Bract.
A reduced leaf or scale-like structure subtending a flower or a grass spikelet in its axil.
2020 International Rules for Seed Testing, (ISTA)
Bract.
In angiosperms: A small or rudimentary leaf or leaf-like structure near the base of a flower or inflorescence.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Bract.
A modified leaf associated with a flower or inflorescence.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
floret, florets
Floret.
General: An individual flower within a cluster. In grasses: A flower usually enclosed by two bracts (lemma and palea).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Floret.
A flower within an inflorescence or in a grass spikelet.
SCST Seed Technologist Training Manual, 2018
Floret.
The lemma and palea with enclosed pistil and stamens, or, in
Poaceae
, the mature caryopsis; for the purpose of the Rules, the term floret refers to the fertile floret, with or without additional sterile lemmas.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Floret.
The lemma and palea with enclosed pistil and stamens or the mature caryopsis in Poaceae; for the purpose of the Rules, the term floret refers to the fertile floret with or without additional sterile lemmas.
2020
International Rules for Seed Testing (ISTA)
Floret.
In grasses, a flower consisting of lemma and palea which enclose the flowering parts -- stamens, pistil and lodicules. May be perfect, staminate, pistillate, neuter, sterile, and so on. A small flower in other plant families.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
palea
Palea.
The upper (inner) bract of a grass floret, sometimes called the inner or upper palea. Bract enclosing the caryopsis on the inner ventral side.
2020
International Rules for Seed Testing (ISTA)
Palea.
The upper (inner) bract of a grass floret, the bract enclosing the caryopsis on the inner (ventral) side.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Palea.
The uppermost of two bracts subtending a grass flower.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Palea.
The upper of the two bracts enclosing the flower in the spikelet of grasses.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
caryopsis
Caryopsis.
In grasses: A dry, one-seeded, indehiscent fruit with the pericarp fused to the seed coat (testa).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Caryopsis.
Naked grass-fruit in which the testa is united with the pericarp.
2020
International Rules for Seed Testing (ISTA)
Caryopsis.
A naked grass fruit in which the testa is fused with the pericarp; also a grain.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
Caryopsis.
The fruit of a grass, which is dry, one-seeded, indehiscent, with the testa and the pericarp completely united.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
ventral
Ventral.
The side facing towards the axis; the lower surface (dorsal).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Ventral.
The lower or front side; opposite to dorsal.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
spikelet, spikelets
Spikelet.
The unit of a grass inflorescence comprising one or more florets subtended by one or two sterile glumes. For the purposes of the Rules, the term spikelet includes, as well as a fertile floret, either one or more additional fertile or completely infertile florets, or glumes.
2020
International Rules for Seed Testing (ISTA)
Spikelet.
Part of a grass inflorescence including one or more florets subtended by one or two glumes.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Spikelet.
In grasses: One or more attached florets usually subtended by one or more bracts (glumes).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Spikelet.
The unit of a grass inflorescence that consists of a pair of empty glumes that enclose one of more florets arranged around a rachilla.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Palmate
Palmate.
Having lobes radiating from a common point; refers especially to
leaf
blades.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
leaf, leaves
Leaf.
Lateral organ of the stem.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Panicle
Panicle.
A compound
inflorescence
with a main
axis
which has
branches
, each of which bears
pedicellate
spikelets
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
inflorescence
Inflorescence.
A flower cluster.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Inflorescence.
The portion of the plant adapted for flowering and fruiting.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
axis
Axis.
The central line of any organ or the support of a group of organs; the main stem of an inflorescence, especially of a panicle.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
branch, branches
Branch.
A lateral stem.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
pedicellate
Pedicellate.
Having, or attached by, a pedicel.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
spikelet, spikelets
Spikelet.
The unit of a grass inflorescence comprising one or more florets subtended by one or two sterile glumes. For the purposes of the Rules, the term spikelet includes, as well as a fertile floret, either one or more additional fertile or completely infertile florets, or glumes.
2020
International Rules for Seed Testing (ISTA)
Spikelet.
Part of a grass inflorescence including one or more florets subtended by one or two glumes.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Spikelet.
In grasses: One or more attached florets usually subtended by one or more bracts (glumes).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Spikelet.
The unit of a grass inflorescence that consists of a pair of empty glumes that enclose one of more florets arranged around a rachilla.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Papilla
Papilla.
An epidermal cell forming a conical protuberance like a
minute
hair
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
minute
Minute.
A small measure of time.
TWS
Minute.
A small thing.
TWS
hair, hairs
Hair.
A uni- or multicellular outgrowth of the epidermis.
2020
International Rules for Seed Testing (ISTA)
Hair.
An elongated uni- or multicellular outgrowth of the epidermis (e.g. in
Anemone
).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Papillionaceous
Papillionaceous.
Butterfly-shaped, as the flower of some
legumes
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
legume, legumes
Legume.
A dry fruit consisting of one carpel, splitting by two longitudinal sutures with a row of seeds on the inner side of the central suture; pod, as in
Fabaceae
.
SCST Seed Technologist Training Manual, 2018
Papillose
Papillose.
Covered with, or bearing, papillae.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Pappus
Pappus.
A ring of fine, sometimes feathery
hairs
or scales, crowning an
achene
.
2020
International Rules for Seed Testing (ISTA)
Pappus.
A ring of fine, sometimes feathery hairs or scales, crowning an achene in
Asteraceae
.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Pappus.
The
teeth
,
awns
, etc., surmounting the achene of the
Asteraceae
family.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
hair, hairs
Hair.
A uni- or multicellular outgrowth of the epidermis.
2020
International Rules for Seed Testing (ISTA)
Hair.
An elongated uni- or multicellular outgrowth of the epidermis (e.g. in
Anemone
).
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
achene, achenium
Achene, achenium.
A dry, indehiscent, one-seeded fruit, formed from one free carpel (e.g.
Ranunculaceae
,
Geum
) with the seed coat distinct from the fruit coat; occasionally consisting of more than one carpel (
Asteraceae
).
2020 International Rules for Seed Testing (ISTA)
Achene, achenium.
A dry, indehiscent, one-seeded fruit, formed strictly from one free carpel, and with the testa distinct from the fruit wall, e.g.
Ranunculaceae
,
Geum
, occasionally consisting of more that one carpel, e.g.
Asteraceae
.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
Achene.
A dry, one-chambered, one-seeded indehiscent fruit with the seed attached to the fruit wall at a single point.
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Achene.
A one-celled, dry indehiscent fruit in which the testa and pericarp are not firmly attached.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
teeth
Teeth.
Pointed lobes or divisions.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
awn, awns
Awn, arista.
Slender, straight or bent bristle. In grasses: usually a continuation of the mid-nerve of lemmas or glumes.
2020 International Rules for Seed Testing (ISTA)
Awn.
The slender bristle extending from the tip or back of the lemma or the glume of a grass spikelet.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil anad Crop Sciences, Colorado State University. Pages 104-11
Pectinate
Pectinate.
Having
teeth
-like projections, giving a comb-like appearance.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
teeth
Teeth.
Pointed lobes or divisions.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Pedicel
Pedicel.
The
stalk
of each single flower in an
inflorescence
.
2020
International Rules for Seed Testing (ISTA)
Pedicel.
A small stalk, the stalk of a single flower in an inflorescence.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Pedicel.
The stalk of a
floret
in an inflorescence or of a grass
spikelet
.
As
sociation of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Pedicel.
The stalk of an individual flower or spikelet of an inflorescence.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
stalk, stalks
Stalk.
The stem of any plant organ.
2020
International Rules for Seed Testing (ISTA)
inflorescence
Inflorescence.
A flower cluster.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Inflorescence.
The portion of the plant adapted for flowering and fruiting.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
floret, florets
Floret.
General: An individual flower within a cluster. In grasses: A flower usually enclosed by two bracts (lemma and palea).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Floret.
A flower within an inflorescence or in a grass spikelet.
SCST Seed Technologist Training Manual, 2018
Floret.
The lemma and palea with enclosed pistil and stamens, or, in
Poaceae
, the mature caryopsis; for the purpose of the Rules, the term floret refers to the fertile floret, with or without additional sterile lemmas.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Floret.
The lemma and palea with enclosed pistil and stamens or the mature caryopsis in Poaceae; for the purpose of the Rules, the term floret refers to the fertile floret with or without additional sterile lemmas.
2020
International Rules for Seed Testing (ISTA)
Floret.
In grasses, a flower consisting of lemma and palea which enclose the flowering parts -- stamens, pistil and lodicules. May be perfect, staminate, pistillate, neuter, sterile, and so on. A small flower in other plant families.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
spikelet, spikelets
Spikelet.
The unit of a grass inflorescence comprising one or more florets subtended by one or two sterile glumes. For the purposes of the Rules, the term spikelet includes, as well as a fertile floret, either one or more additional fertile or completely infertile florets, or glumes.
2020
International Rules for Seed Testing (ISTA)
Spikelet.
Part of a grass inflorescence including one or more florets subtended by one or two glumes.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Spikelet.
In grasses: One or more attached florets usually subtended by one or more bracts (glumes).
Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Spikelet.
The unit of a grass inflorescence that consists of a pair of empty glumes that enclose one of more florets arranged around a rachilla.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Pedicellate
Pedicellate.
Having, or attached by, a
pedicel
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
pedicel
Pedicel.
The stalk of each single flower in an inflorescence.
2020
International Rules for Seed Testing (ISTA)
Pedicel.
A small stalk, the stalk of a single flower in an inflorescence.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Pedicel.
The stalk of a floret in an inflorescence or of a grass spikelet.
As
sociation of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
Pedicel.
The stalk of an individual flower or spikelet of an inflorescence.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Peduncle
Peduncle.
The
stalk
or stem bearing an
inflorescence
.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
stalk, stalks
Stalk.
The stem of any plant organ.
2020
International Rules for Seed Testing (ISTA)
inflorescence
Inflorescence.
A flower cluster.
ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
Inflorescence.
The portion of the plant adapted for flowering and fruiting.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
Pendant
Pendant.
Hanging down.
Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
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