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Germination

  • Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)

  • Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

germination

  • Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)

  • Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

ISTA

International Rules for Seed Testing

seedling, seedlings

  • Seedling. A young plant developing from the embryo of a seed. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

essential structure, essential structures

  • Essential structure. Structure which is critical for continued development of the seedling into a plant. SCST Seed Technologist Training Manual. 2018

  • Essential structure. Any seedling structure that must be considered when classifying a seedling as either normal or abnormal. In general, an essential structure is one that is critical for successful establishment and development of a seedling into a plant. Structure which is critical for continued development of the seedling into a plant. Baalbaki, R.Z. Germination & Dormancy. TWS Website. 2020

embryo, embryos

  • Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)

  • Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

kind , kind of seed

  • Kind (of seed). One or more related species or subspecies that singly or collectively is usually known by one common name. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

normal seedling, normal

  • Normal seedling. A seedling with all essential structures present and capable of developing into a plant under favorable conditions; certain defects may be present if they are judged to be not so severe as to impede further development of the plant (see abnormal seedling). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

radicle, radicles

  • Radicle. The initial root of the embryo, which develops into the primary root after emergence through the testa during germination. ISTA Handbook of Pure Seed Definitions, Glossary, 3rd Edition, 2010

  • Radicle. The part of an embro giving rise to the root system of a plant. Compare to plumule. SCST Seed Technologist Training Manual, 2018

  • Radicle. The lower portion of the hypocotyl which grows into the primary root. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

  •  Radicles. Spring from the root; clustered at the base of the stem. TWS

Germination working sample

  • Germination working sample. The sub-sample taken either from the pure seed portion of the purity analysis or directly from the submitted sample on which the germination test is performed. Refer to sections 2.3 c and 6.1. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

purity analysis

  • Purity analysis. The objective of a purity analysis is to determine the physical composition of the working sample. The analysis shall include the identification of the kind, or kind and cultivar of seed under consideration, and all contaminating species and inert matter. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Purity. The term “purity” means the name or names of the kind, type, or variety and the percentage or percentages thereof; the percentage of other agricultural seed or crop seed; the percentage of weed seeds, including noxious-weeds seeds; the percentage of inert matter; and the names of the noxious-weed seeds and the rate of occurrence of each. Agricultural Marketing Service, USDA. Federal Seed Act, Part 201. Federal Seed Act Regulations. 201.2 Terms Defined. Current as of May 21, 2020.

germination

  • Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)

  • Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

Gibberellic acid

  • Gibberellic acid. A growth hormone, one of over 50 gibberellins. First discovered in the fungus Gibberella fujikuroi. Found in highest concentrations in immature seeds. Can be used to substitute for dormancy-breaking cold and light requirements in many species. SCST Seed Technologist Training Manual, 2018

growth hormone

  • Growth hormone. A chemical compound generally produced in one part of an organism and transported to another part of the organism where it controls and affects growth and development. SCST Seed Technologist Training Manual, 2018

gibberellins

  • Gibberellins. Growth hormones that stimulate cell division and cell elongation. SCST Seed Technologist Training Manual, 2018

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

dormant seeds, dormancy

  • Dormant seeds. Viable seeds, other than hard seeds, which fail to germinate when provided the specified germination conditions for the kind of seed in question. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Dormancy. Delayed germination or growth; a condition of inactivity. SCST Seed Technologist Training Manual, 2018

species

  • Species. A category of classification lower than a genus that is made up of plants which possess in common distinctive characteristics that are reproduced in their offspring. The species name included second to the genus in the scientific binomial. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Gibberellins

  • Gibberellins. Growth hormones that stimulate cell division and cell elongation. SCST Seed Technologist Training Manual, 2018

Glabrous

  • Glabrous. Smooth, without hairs. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

hair, hairs

  • Hair. A uni- or multicellular outgrowth of the epidermis. 2020 International Rules for Seed Testing (ISTA)

  • Hair. An elongated uni- or multicellular outgrowth of the epidermis (e.g. in Anemone). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

Glandular

  • Glandular. Bears organs for secreting a substance or substances. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Glaucous

  • Gloucous. Covered with a powdery wax coating such as found on a Concord grape. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Globose

  • Globose. Shape is spherical or nearly so. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Glume, glumes

  • Glume. In grasses: A bract—often paired—at the base of a spikelet. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Glume. One of the two usually sterile bracts at the base of a grass spikelet. 2020 International Rules for Seed Testing (ISTA)

  • Glumes. The pair of empty bracts at the base of a spikelet. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

bract

  • Bract. A reduced leaf or scale-like structure subtending a flower or a grass spikelet in its axil. 2020 International Rules for Seed Testing, (ISTA)

  • Bract. In angiosperms: A small or rudimentary leaf or leaf-like structure near the base of a flower or inflorescence. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Bract. A modified leaf associated with a flower or inflorescence. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

spikelet, spikelets

  • Spikelet. The unit of a grass inflorescence comprising one or more florets subtended by one or two sterile glumes. For the purposes of the Rules, the term spikelet includes, as well as a fertile floret, either one or more additional fertile or completely infertile florets, or glumes. 2020 International Rules for Seed Testing (ISTA)

  • Spikelet. Part of a grass inflorescence including one or more florets subtended by one or two glumes. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Spikelet. In grasses: One or more attached florets usually subtended by one or more bracts (glumes). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Spikelet. The unit of a grass inflorescence that consists of a pair of empty glumes that enclose one of more florets arranged around a rachilla. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

sterile

  • Sterile. Without functional sex organs (for grass florets: without caryopsis). 2020 International Rules for Seed Testing (ISTA)

  • Sterile. Without pistils. A sterile floret may be staminate or neuter. It may even lack a palea, and consist of nothing but a lemma. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Grain

  • Grain. The seed-like fruit of any cereal. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

fruit

  • Fruit. The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Fruit. In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity. SCST Seed Technologist Training Manual, 2018

  • Fruit. The ripened ovary of a seed plant and associated parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Gravimetric

  • Gravimetric. Determined by weighing. Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020

Gregarious

  • Gregarious. Growing in groups or colonies. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Growth hormone

  • Growth hormone. A chemical compound generally produced in one part of an organism and transported to another part of the organism where it controls and affects growth and development. SCST Seed Technologist Training Manual, 2018

Gynoecium

  • Gynoecium. The female reproductive part of a flower, composed of one or more pistils. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Gynoecium. The carpels taken collectively. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

carpel, carpels

  • Carpel. The female reproductive organ of flowering plants. It consists of an ovary, containing one or more ovules (which became seeds after fertilization), and a stigma, a surface receptive for pollen grains. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Carpel. A simple pistil or an element of a compound pistil. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

H

Hair, hairs

  • Hair. A uni- or multicellular outgrowth of the epidermis. 2020 International Rules for Seed Testing (ISTA)

  • Hair. An elongated uni- or multicellular outgrowth of the epidermis (e.g. in Anemone). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

epidermis

  • Epidermis. The surface layer of cells of leaves and other soft plant parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Hairy

  • Hairy. Bearing hairs. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

hair, hairs

  • Hair. A uni- or multicellular outgrowth of the epidermis. 2020 International Rules for Seed Testing (ISTA)

  • Hair. An elongated uni- or multicellular outgrowth of the epidermis (e.g. in Anemone). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

Hard seed, hard seeds

  • Hard seed. Seeds which remain hard at the end of the prescribed test period because they have not absorbed water due to an impermeable seed coat. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Head, heads

  • Head. A type of inflorescence with sessile or nearly sessile flowers on a very short axis or receptacle, as in a sunflower. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

inflorescence

  • Inflorescence. A flower cluster. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Inflorescence. The portion of the plant adapted for flowering and fruiting. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

sessile

  • Sessile. Without a stalk or pedicel. 2020 International Rules for Seed Testing (ISTA)

  • Sessile. Attached directly by the base and not raised on a stalk. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

axis

  • Axis. The central line of any organ or the support of a group of organs; the main stem of an inflorescence, especially of a panicle. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

receptacle

  • Receptacle. Apex of a pedicel or peduncle from which the organs of a flower arise. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Herbaceous

  • Herbaceous. Describing plants which do not develop much woody tissue but remain soft and succulent. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

succulent

  • Succulent. Fruits in which the mesocarp develops into a fleshy covering. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Succulent. Fleshy or juicy. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Hesperidium

  • Hesperidium. A type of berry with a leathery epicarp, such as a citrus fruit. Fluid filled trichomes fill the locule of each carpel to form the characteristic segments. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

berry

  • Berry. A many seeded fleshy indehiscent fruit. The pericarp usually forms a tough outer skin and the mesocarp becomes massive and fleshy. The epicarp and mesocarp may be highly coloured to attract the animals that act as agents of dispersal.  ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.

epicarp

  • Epicarp. The outer layer of the pericarp. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

fruit

  • Fruit. The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Fruit. In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity. SCST Seed Technologist Training Manual, 2018

  • Fruit. The ripened ovary of a seed plant and associated parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

locule, loculus

  • Locule, loculus, loculi (plural). Compartment of the ovary containing the seeds. 2020 International Rules for Seed Testing (ISTA)

  • Locule, loculus, loculi (plural). A compartment of the ovary containing the ovules, and later, seeds. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Locule, loculus, loculi (plural). One of the cavities of an ovary. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

carpel, carpels

  • Carpel. The female reproductive organ of flowering plants. It consists of an ovary, containing one or more ovules (which became seeds after fertilization), and a stigma, a surface receptive for pollen grains. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Carpel. A simple pistil or an element of a compound pistil. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Heterogeneous

  • Heterogeneous. Having unlike qualities. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Heterogeneous seed sample of lot, heterogeneous

  • Heterogeneous seed sample or lot. A sample or lot which is not uniform throughout in one or more characteristics. SCST Seed Technologist Training Manual, 2018

Hilum

  • Hilum. Scar on a seed coat marking the place of attachment of the seed stalk to the seed. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
 

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

stalk, stalks

  • Stalk. The stem of any plant organ. 2020 International Rules for Seed Testing (ISTA)

Hirsute

  • Hirsute. Pubescent with rather straight stiff hairs. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

pubescent

  • Pubescent. Covered with soft hairs. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

hair, hairs

  • Hair. A uni- or multicellular outgrowth of the epidermis. 2020 International Rules for Seed Testing (ISTA)

  • Hair. An elongated uni- or multicellular outgrowth of the epidermis (e.g. in Anemone). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

Hispid

  • Hispid. Pubescent with stiff or rigid hairs. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

pubescent

  • Pubescent. Covered with soft hairs. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

rigid

  • Rigid. Stiff, not flexible. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

hair, hairs

  • Hair. A uni- or multicellular outgrowth of the epidermis. 2020 International Rules for Seed Testing (ISTA)

  • Hair. An elongated uni- or multicellular outgrowth of the epidermis (e.g. in Anemone). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

Histo-differentiation

  • Histo-differentiation. The differentiation of cells in an embryo. Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020

differentiation

  • Differentiation. The process by which different cell types are formed. Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020

embryo, embryos

  • Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)

  • Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Homogeneous

  • Homogeneous. A sample or seed lot which is uniform throughout. SCST Seed Technologist Training Manual, 2018

seed lot

  • Seed lot. A seed lot is a specified quantity of seed that is physically and uniquely identified. 2020 International Rules for Seed Testing, (ISTA)

  • Lot of seed. The  term "lot of seed" means a definite quantity of seed identified by a number, every portion or bag which is uniform, within permitted tolerances, for the factors which appear in the labeling. Agricultural Marketing Service, USDA. Federal Seed Act, Part 201. Federal Seed Act Regulations. 201.2 Terms Defined. Current as of May 21, 2020.

Hulled seed

  • Hulled seed. Seed units ordinarily with various structures, including but not limited to wings, hairs, spines, awns, sterile florets, calyces, etc., attached to or surrounding the seed (or caryopsis) that have these structures partially or completely removed during seed conditioning to improve seed flow and plantability. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

wing, wings

  • Wing. A flat membranous outgrowth from a fruit or seed. 2020 International Rules for Seed Testing (ISTA)

  • Wing. A flat membranous outgrowth from a fruit or seed (e.g. Coreopsis, Dimorphotheca). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Wing. Lateral petal of a papilionaceous corolla, as in some legumes. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

hair, hairs

  • Hair. A uni- or multicellular outgrowth of the epidermis. 2020 International Rules for Seed Testing (ISTA)

  • Hair. An elongated uni- or multicellular outgrowth of the epidermis (e.g. in Anemone). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

awn, awns

  • Awn, arista. Slender, straight or bent bristle. In grasses: usually a continuation of the mid-nerve of lemmas or glumes. 2020 International Rules for Seed Testing (ISTA)

  • Awn. The slender bristle extending from the tip or back of the lemma or the glume of a grass spikelet. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil anad Crop Sciences, Colorado State University. Pages 104-11

sterile floret, sterile florets

  • Sterile floret. A floret with no caryopsis. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

calyx, calyces

  • Calyx, calyces (plural). The outer floral envelope composed of the sepals. 2020 International Rules for Seed Testing (ISTA)

  • Calyx, calyces (plural). The outer floral envelope of dicotyledons, composed of the sepals. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.

  • Calyx. The outer cycle of the perianth; the sepals considered collectively. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

caryopsis

  • Caryopsis. In grasses: A dry, one-seeded, indehiscent fruit with the pericarp fused to the seed coat (testa). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Caryopsis. Naked grass-fruit in which the testa is united with the pericarp. 2020 International Rules for Seed Testing (ISTA)

  • Caryopsis. A naked grass fruit in which the testa is fused with the pericarp; also a grain. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.

  • Caryopsis. The fruit of a grass, which is dry, one-seeded, indehiscent, with the testa and the pericarp completely united. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Hyaline

  • Hyaline. Thin and translucent or transparent. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

translucent

  • Translucent. Permits some passage of light, but diffuses it. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Hydration

  • Hydration. The act of becoming chemically combined with water. SCST Seed Technologist Training Manual, 2018

Hydrolyzed

  • Hydrolyzed. Chemically decomposed by taking up the elements of water. SCST Seed Technologist Training Manual, 2018

Hygroscopic

  • Hygroscopic. The phenomenon where seeds will take up or lose water until they have reached the same relative humidity as that of their environment. Bowden, L., Moisture Testing in Wild Species, TWS Website, 2020.

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Hypanthium, hypanthia

  • Hypanthiumhypanthia (plural). A ring-like, cup-like or tubular structure which surrounds the ovary and on which sepals, petals and stamens are borne. 2020 International Rules for Seed Testing (ISTA)

  • Hypanthium. A cup-shaped structure surrounding the ovary, derived either from the fusion of floral parts or an extension of the receptacle to which the floral parts are attached. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

ovary

  • Ovary. The basal, expanded part of the pistil that contains the ovules. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Ovary. The basal enlarged portion of a pistil within which seeds develop. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

petal, petals

  • Petal. One of the divisions on a corolla; a showy floral leaf of the perianth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

receptacle

  • Receptacle. Apex of a pedicel or peduncle from which the organs of a flower arise. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Hypocotyl

  • Hypocotyl. The portion of the seedling axis between the root and the cotyledons is the hypocotyl. The hypocotyl is a transition structure for the transport of water and dissolved salts from the roots to the epicotyl. When a seed with epigeal germination (see section 2.5 for definitions of epigeal and hypogeal germination) is planted in moist soil, the hypocotyl elongates carrying the cotyledons above the soil surface.

    In monocotyledons the hypocotyl is usually not discernible as a separate structure. The mesocotyl is the part of the seedling axis between the scutellum and the base of the coleoptile. In some species (e.g. Zea mays) the elongation of the mesocotyl may be considerable. In others (e.g. Triticum aestivum) the elongation may be imperceptible. Elongation of the mesocotyl is suppressed by light after the coleoptile emerges from the soil.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Hypocotyl. That portion of the embryo stem below the cotyledons. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

seedling, seedlings

  • Seedling. A young plant developing from the embryo of a seed. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

axis

  • Axis. The central line of any organ or the support of a group of organs; the main stem of an inflorescence, especially of a panicle. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

root, roots

  • Root. In dicotyledons and gymnosperms, the root system serves three major functions: (1) to anchor the plant in the soil, (2) to absorb water and dissolved salts from the soil and (3) to conduct the water and salts to the hypocotyl, cotyledons and epicotyl. The embryonic root, or radicle, is located at the basal end of the embryo and is usually the first seedling structure to rupture the testa. After emergence it is referred to as the primary root. The primary root elongates rapidly and soon numerous root hairs develop, greatly increasing the absorbing surface of the roots. As the seedling continues to grow, secondary roots develop from the primary root and from other secondary roots. Roots may also emerge from other structures (e.g. the hypocotyl) and are referred to as adventitious roots. As in the dicotyledons, the monocotyledon root system serves to anchor the plant in soil, absorb water and dissolved salts from the soil and to conduct the water and salts to the growing seedling. The embryonic root, or radicle, is situated at the basal end of the embryo and, in the case of the Poaceae, its apex is covered by the coleorhiza. After the radicle emerges it is referred to as the primary root. In some species of the Poaceae (e.g. Triticum) the primary root is indistinguishable from the other roots that develop from the scutellar node region and hence all of these are referred to as seminal roots. Roots that develop from structures above the scutellar or cotyledonary node are called adventitious roots. Secondary roots may develop from seminal and adventitious roots.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Root. A portion of a higher plant bearing neither leaves nor reproductive organs, usually underground. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

cotyledon, cotyledons

  • Cotyledon. The first leaf or pair of leaves of an embryo and seedling, often a food-storage organ. The cotyledons may remain in the seed (hypogeal germination) or emerge to become the first photosynthetic organs (epigeal germination). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.

  • Cotyledons. The cotyledons are the storage structures of the embryo. They may be only a small portion of the seed in species with endosperm, perisperm or female gametophyte storage tissue, or they may occupy a large portion of the embryo when they are the primary storage tissue (e.g. Phaseolus vulgaris). In epigeal species, the cotyledons may grow quite large and become the first photosynthetic structures of the young plant. In hypogeal species the primary function of the cotyledons is to provide nutrients to the growing seedling until it can produce its own nutrients. In most species the cotyledons shrivel and drop off as their reserves are depleted. In a few species (e.g. Cucurbita pepo, pumpkin) the cotyledons may persist well beyond the seedling stage of growth. In the monocotyledons the cotyledon absorbs nutrients from the endosperm and transfers them to the growing seedling. In the Poaceae the cotyledon is called the scutellum. It is in close proximity to the endosperm and is laterally attached to the embryo axis. In Allium (Liliaceae) the cotyledon tip remains embedded in the endosperm to absorb nutrients but the cotyledon also emerges from the soil (i.e. germination is epigeal) and becomes photosynthetic. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Cotyledon. The first leaves of the embryo that serve for food digestion and food storage. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

hypocotyl

  • Hypocotyl. The portion of the seedling axis between the root and the cotyledons is the hypocotyl. The hypocotyl is a transition structure for the transport of water and dissolved salts from the roots to the epicotyl. When a seed with epigeal germination (see section 2.5 for definitions of epigeal and hypogeal germination) is planted in moist soil, the hypocotyl elongates carrying the cotyledons above the soil surface.

    In monocotyledons the hypocotyl is usually not discernible as a separate structure. The mesocotyl is the part of the seedling axis between the scutellum and the base of the coleoptile. In some species (e.g. Zea mays) the elongation of the mesocotyl may be considerable. In others (e.g. Triticum aestivum) the elongation may be imperceptible. Elongation of the mesocotyl is suppressed by light after the coleoptile emerges from the soil.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Hypocotyl. That portion of the embryo stem below the cotyledons. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

root, roots

  • Root. In dicotyledons and gymnosperms, the root system serves three major functions: (1) to anchor the plant in the soil, (2) to absorb water and dissolved salts from the soil and (3) to conduct the water and salts to the hypocotyl, cotyledons and epicotyl. The embryonic root, or radicle, is located at the basal end of the embryo and is usually the first seedling structure to rupture the testa. After emergence it is referred to as the primary root. The primary root elongates rapidly and soon numerous root hairs develop, greatly increasing the absorbing surface of the roots. As the seedling continues to grow, secondary roots develop from the primary root and from other secondary roots. Roots may also emerge from other structures (e.g. the hypocotyl) and are referred to as adventitious roots. As in the dicotyledons, the monocotyledon root system serves to anchor the plant in soil, absorb water and dissolved salts from the soil and to conduct the water and salts to the growing seedling. The embryonic root, or radicle, is situated at the basal end of the embryo and, in the case of the Poaceae, its apex is covered by the coleorhiza. After the radicle emerges it is referred to as the primary root. In some species of the Poaceae (e.g. Triticum) the primary root is indistinguishable from the other roots that develop from the scutellar node region and hence all of these are referred to as seminal roots. Roots that develop from structures above the scutellar or cotyledonary node are called adventitious roots. Secondary roots may develop from seminal and adventitious roots.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Root. A portion of a higher plant bearing neither leaves nor reproductive organs, usually underground. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

epicotyl, epicotyls

  • Epicotyl. In Dicotyledons and gymnosperms, the epicotyl includes all seedling structures above the cotyledons. In species with epigeal germination (e.g. Phaseolus vulgaris), the epicotyl, cotyledons and part of the hypocotyl emerge from the soil. In species with hypogeal germination (e.g. Pisum sativum), only the epicotyl emerges, carrying the first foliage leaves above the soil surface. In these species, the epicotyl also bears one or more scale leaves. Dormant meristematic buds in the axils of these scale leaves become active if there is damage to the terminal bud. The conducting tissue of the epicotyl transfers water and nutrients from the hypocotyl and cotyledons to the leaves and terminal bud above. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Epicotyl. The stem of the embryo or young seedling above the cotyledons. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

epigeal, epigeal germination

  • Epigeal. Cotyledons borne above the ground after germination (see hypogeal). Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

  • Epigeal germination. A type of germination in which cotyledons are carried above soil level by the elongating hypocotyl (see hypogeal germination). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

epigeal, epigeal germination

  • Epigeal. Cotyledons borne above the ground after germination (see hypogeal). Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

  • Epigeal germination. A type of germination in which cotyledons are carried above soil level by the elongating hypocotyl (see hypogeal germination). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

hypogeal

  • Hypogeal. The cotyledons borne below the ground after germination (see epigeal). Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

germination

  • Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)

  • Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

monocotyledon, monocotyledons

  • Monocotyledon. Plant having one cotyledon, as in the grass. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

mesocotyl

  • Mesocotyl. A term applied to the sub-crown internode of a grass seedling. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

scutellum

  • Scutellum. Shield-shaped organ surrounding the embryo of a grass that is morphologically like a cotyledon. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

coleoptile, coleoptiles

  • Coleoptile. The coleoptile is only present in species of the Poaceae. It is a leaf-like, cylindrical sheath enclosing the terminal bud of the embryo and the developing leaves of the young seedling. The coleoptile provides protection for the leaves as they push up through the soil. After emergence from the soil, growth of the leaves ordinarily causes the coleoptile to split downward from the tip. The coleoptile does not persist beyond the seedling stage. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Coleoptile. The sheath covering the first leaf of a grass seedling. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

species

  • Species. A category of classification lower than a genus that is made up of plants which possess in common distinctive characteristics that are reproduced in their offspring. The species name included second to the genus in the scientific binomial. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

embryo, embryos

  • Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)

  • Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Hypogeal

  • Hypogeal. The cotyledons borne below the ground after germination (see epigeal). Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

cotyledon, cotyledons

  • Cotyledon. The first leaf or pair of leaves of an embryo and seedling, often a food-storage organ. The cotyledons may remain in the seed (hypogeal germination) or emerge to become the first photosynthetic organs (epigeal germination). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.

  • Cotyledons. The cotyledons are the storage structures of the embryo. They may be only a small portion of the seed in species with endosperm, perisperm or female gametophyte storage tissue, or they may occupy a large portion of the embryo when they are the primary storage tissue (e.g. Phaseolus vulgaris). In epigeal species, the cotyledons may grow quite large and become the first photosynthetic structures of the young plant. In hypogeal species the primary function of the cotyledons is to provide nutrients to the growing seedling until it can produce its own nutrients. In most species the cotyledons shrivel and drop off as their reserves are depleted. In a few species (e.g. Cucurbita pepo, pumpkin) the cotyledons may persist well beyond the seedling stage of growth. In the monocotyledons the cotyledon absorbs nutrients from the endosperm and transfers them to the growing seedling. In the Poaceae the cotyledon is called the scutellum. It is in close proximity to the endosperm and is laterally attached to the embryo axis. In Allium (Liliaceae) the cotyledon tip remains embedded in the endosperm to absorb nutrients but the cotyledon also emerges from the soil (i.e. germination is epigeal) and becomes photosynthetic. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Cotyledon. The first leaves of the embryo that serve for food digestion and food storage. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

germination

  • Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)

  • Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

epigeal, epigeal germination

  • Epigeal. Cotyledons borne above the ground after germination (see hypogeal). Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

  • Epigeal germination. A type of germination in which cotyledons are carried above soil level by the elongating hypocotyl (see hypogeal germination). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

Hypogynous

  • Hypogynous. A flower with sepals, petals, and stamens attached to the receptacle below the ovary (superior ovary). SCST Seed Technologist Training Manual, 2018

petal, petals

  • Petal. One of the divisions on a corolla; a showy floral leaf of the perianth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

receptacle

  • Receptacle. Apex of a pedicel or peduncle from which the organs of a flower arise. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

ovary

  • Ovary. The basal, expanded part of the pistil that contains the ovules. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Ovary. The basal enlarged portion of a pistil within which seeds develop. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Hysteresis

  • Hysteresis. The effect where at a given relative humidity seeds will have a different moisture content dependent on whether they are absorbing (gaining) or desorbing (losing) moisture. Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

I

Imbibition

  • Imbibition. The uptake of water by the seed from the germination substrate. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

germination

  • Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)

  • Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

Imbricate

  • Imbricate. Overlapping like shingles on a roof. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Impaired

  • Impaired. Unable to function normally, in reference to damaged seedling structures. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

seedling, seedlings

  • Seedling. A young plant developing from the embryo of a seed. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

Incise

  • Incise. Cut deeply. TWS

Indehiscent

  • Indehiscent. Not opening; fruits which do not open at maturity. 2020 International Rules for Seed Testing (ISTA)

  • Indehiscent. Fruits in which the fruit wall does not split open at maturity to release the seed (see dehiscent). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Indehiscent fruit. A fruit that does not open at maturity. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

  • Indehiscent. Fruits remaining closed at maturity. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

open

  • Open. Loose; opposite of dense or compact. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

fruit

  • Fruit. The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Fruit. In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity. SCST Seed Technologist Training Manual, 2018

  • Fruit. The ripened ovary of a seed plant and associated parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

dehiscent fruit, dehiscent

  • Dehiscent fruit. A dry fruit, that experiences desiccation. Seeds within dehiscent fruits will lose moisture content during the later stages of seed development. Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020

  • Dehiscent fruit. A fruit that opens at maturity allowing seeds to be released from the fruit. SCST Seed Technologist Training Manual, 2018

  • Dehiscent. A fruit in which the fruit wall splits open at maturity to release the seed. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Dehiscent. Opening spontaneously at maturity. SCST Seed Technologist Training Manual, 2018

Indigenous

  • Indigenous. Native to the region of growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Indurate

  • Indurate. Hardened. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Inert matter, inert

  • Inert matter.  Inert matter shall include seeds and seed-like structures from both crop and weed plants and other materials not described in section 3.2 and Table 3A or can be described as follows: seeds and seed-like structure from crop and weed plants and other non-seed matter. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

inert matter

  • Inert matter.  Inert matter shall include seeds and seed-like structures from both crop and weed plants and other materials not described in section 3.2 and Table 3A or can be described as follows: seeds and seed-like structure from crop and weed plants and other non-seed matter. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

seed, seeds

  • Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

  • Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Infection

  • Infection. Entrance and spread of disease organisms in living material (e.g., seedling structures) often causing disease symptoms and decay. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

seedling, seedlings

  • Seedling. A young plant developing from the embryo of a seed. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

decay

  • Decay. Break-down of organic tissue, usually associated with the presence of microorganisms. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

Inferior ovary

  • Inferior ovary. An ovary completely or partially surrounded by floral parts of embedded in receptacle tissue. SCST Seed Technologist Training Manual, 2018

ovary

  • Ovary. The basal, expanded part of the pistil that contains the ovules. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Ovary. The basal enlarged portion of a pistil within which seeds develop. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

receptacle

  • Receptacle. Apex of a pedicel or peduncle from which the organs of a flower arise. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Inflated

  • Inflated. Puffed up, bladdery. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Inflexed

  • Inflexed. Turned in at the margins. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

Inflorescence

  • Inflorescence. A flower cluster. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

  • Inflorescence. The portion of the plant adapted for flowering and fruiting. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

cluster

  • Cluster. A densely crowded inflorescence or, in Beta, part of an inflorescence. 2020 International Rules for Seed Testing (ISTA)