• Elliptical. Shaped like an ellipse. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Rudimentary. Imperfectly developed. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Axis. The central line of any organ or the support of a group of organs; the main stem of an inflorescence, especially of a panicle. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Cotyledon. The first leaf or pair of leaves of an embryo and seedling, often a food-storage organ. The cotyledons may remain in the seed (hypogeal germination) or emerge to become the first photosynthetic organs (epigeal germination). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Cotyledons. The cotyledons are the storage structures of the embryo. They may be only a small portion of the seed in species with endosperm, perisperm or female gametophyte storage tissue, or they may occupy a large portion of the embryo when they are the primary storage tissue (e.g. Phaseolus vulgaris). In epigeal species, the cotyledons may grow quite large and become the first photosynthetic structures of the young plant. In hypogeal species the primary function of the cotyledons is to provide nutrients to the growing seedling until it can produce its own nutrients. In most species the cotyledons shrivel and drop off as their reserves are depleted. In a few species (e.g. Cucurbita pepo, pumpkin) the cotyledons may persist well beyond the seedling stage of growth. In the monocotyledons the cotyledon absorbs nutrients from the endosperm and transfers them to the growing seedling. In the Poaceae the cotyledon is called the scutellum. It is in close proximity to the endosperm and is laterally attached to the embryo axis. In Allium (Liliaceae) the cotyledon tip remains embedded in the endosperm to absorb nutrients but the cotyledon also emerges from the soil (i.e. germination is epigeal) and becomes photosynthetic. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Cotyledon. The first leaves of the embryo that serve for food digestion and food storage. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Angiosperm. A plant whose seeds are borne within a mature ovary (fruit). SCST Seed Technologist Training Manual, 2018

• Fertilization. A sexual process in which two dissimilar gametes fuse. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)
  
  
• Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Endosperm. Nutritive tissue originating from fertilization and retained at maturity in some seeds as a storage tissue for food reserves. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Endosperm.The nutritive tissue developed as a result of fertilization associated with the embryo in seeds. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
  
  
• Endosperm. Endosperm is one of the products of double fertilization, and in some species, particularly those of the Poaceae, it develops as the storage tissue. In this event, little or no nucellus tissue remains, and the extent of cotyledon development varies. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Species. A category of classification lower than a genus that is made up of plants which possess in common distinctive characteristics that are reproduced in their offspring. The species name included second to the genus in the scientific binomial. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Radicle. The initial root of the embryo, which develops into the primary root after emergence through the testa during germination. ISTA Handbook of Pure Seed Definitions, Glossary, 3rd Edition, 2010
  
  
• Radicle. The part of an embro giving rise to the root system of a plant. Compare to plumule. SCST Seed Technologist Training Manual, 2018
  
  
• Radicle. The lower portion of the hypocotyl which grows into the primary root. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
  
  
•  Radicles. Spring from the root; clustered at the base of the stem. TWS

• Hypocotyl. The portion of the seedling axis between the root and the cotyledons is the hypocotyl. The hypocotyl is a transition structure for the transport of water and dissolved salts from the roots to the epicotyl. When a seed with epigeal germination (see section 2.5 for definitions of epigeal and hypogeal germination) is planted in moist soil, the hypocotyl elongates carrying the cotyledons above the soil surface.
  
  In monocotyledons the hypocotyl is usually not discernible as a separate structure. The mesocotyl is the part of the seedling axis between the scutellum and the base of the coleoptile. In some species (e.g. Zea mays) the elongation of the mesocotyl may be considerable. In others (e.g. Triticum aestivum) the elongation may be imperceptible. Elongation of the mesocotyl is suppressed by light after the coleoptile emerges from the soil.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Hypocotyl. That portion of the embryo stem below the cotyledons. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Epicotyl. In Dicotyledons and gymnosperms, the epicotyl includes all seedling structures above the cotyledons. In species with epigeal germination (e.g. Phaseolus vulgaris), the epicotyl, cotyledons and part of the hypocotyl emerge from the soil. In species with hypogeal germination (e.g. Pisum sativum), only the epicotyl emerges, carrying the first foliage leaves above the soil surface. In these species, the epicotyl also bears one or more scale leaves. Dormant meristematic buds in the axils of these scale leaves become active if there is damage to the terminal bud. The conducting tissue of the epicotyl transfers water and nutrients from the hypocotyl and cotyledons to the leaves and terminal bud above. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Epicotyl. The stem of the embryo or young seedling above the cotyledons. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Leaf. Lateral organ of the stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Essential structure. Structure which is critical for continued development of the seedling into a plant. SCST Seed Technologist Training Manual. 2018
  
  
• Essential structure. Any seedling structure that must be considered when classifying a seedling as either normal or abnormal. In general, an essential structure is one that is critical for successful establishment and development of a seedling into a plant. Structure which is critical for continued development of the seedling into a plant. Baalbaki, R.Z. Germination & Dormancy. TWS Website. 2020

• Seedling. A young plant developing from the embryo of a seed. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)
  
  
• Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Dormant seeds. Viable seeds, other than hard seeds, which fail to germinate when provided the specified germination conditions for the kind of seed in question. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Dormancy. Delayed germination or growth; a condition of inactivity. SCST Seed Technologist Training Manual, 2018

• Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)
  
  
• Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

• Viable. Alive. Seed viability indicates that a seed contains structures and substances including enzyme systems that give it the capacity to germinate under favorable conditions in the absence of dormancy.  SCST Seed Technologist Training Manual, 2018 

• Dormant seeds. Viable seeds, other than hard seeds, which fail to germinate when provided the specified germination conditions for the kind of seed in question. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Dormancy. Delayed germination or growth; a condition of inactivity. SCST Seed Technologist Training Manual, 2018

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Ovule. The immature seed within the ovary. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Ovule. A structure, consisting of a female gametophyte, nucellus, and integuments, which develops into a seed after fertilization. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)
  
  
• Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fertilization. A sexual process in which two dissimilar gametes fuse. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Pericarp [fruit coat]. The wall of the mature ovary or fruit. 2020 International Rules for Seed Testing (ISTA)
  
  
• Pericarp. The ovary wall. the fruit coat enclosing the seed. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Pericarp. Fruit wall. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Pericarp. The wall of a ripened ovary; the fruit coat.  Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fruit. The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Fruit. In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity. SCST Seed Technologist Training Manual, 2018
  
  
• Fruit. The ripened ovary of a seed plant and associated parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)
  
  
• Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fertilization. A sexual process in which two dissimilar gametes fuse. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Storage tissue. The storage tissue in seeds may originate from four sources depending on the species: (1) perisperm; (2) endosperm; (3) cotyledons and (4) female gametophyte.   Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)
  
  
• Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Endosperm. Nutritive tissue originating from fertilization and retained at maturity in some seeds as a storage tissue for food reserves. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Endosperm.The nutritive tissue developed as a result of fertilization associated with the embryo in seeds. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
  
  
• Endosperm. Endosperm is one of the products of double fertilization, and in some species, particularly those of the Poaceae, it develops as the storage tissue. In this event, little or no nucellus tissue remains, and the extent of cotyledon development varies. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Species. A category of classification lower than a genus that is made up of plants which possess in common distinctive characteristics that are reproduced in their offspring. The species name included second to the genus in the scientific binomial. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Nucellus. The ovule tissue within the integuments around the embryo sac. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Cotyledon. The first leaf or pair of leaves of an embryo and seedling, often a food-storage organ. The cotyledons may remain in the seed (hypogeal germination) or emerge to become the first photosynthetic organs (epigeal germination). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Cotyledons. The cotyledons are the storage structures of the embryo. They may be only a small portion of the seed in species with endosperm, perisperm or female gametophyte storage tissue, or they may occupy a large portion of the embryo when they are the primary storage tissue (e.g. Phaseolus vulgaris). In epigeal species, the cotyledons may grow quite large and become the first photosynthetic structures of the young plant. In hypogeal species the primary function of the cotyledons is to provide nutrients to the growing seedling until it can produce its own nutrients. In most species the cotyledons shrivel and drop off as their reserves are depleted. In a few species (e.g. Cucurbita pepo, pumpkin) the cotyledons may persist well beyond the seedling stage of growth. In the monocotyledons the cotyledon absorbs nutrients from the endosperm and transfers them to the growing seedling. In the Poaceae the cotyledon is called the scutellum. It is in close proximity to the endosperm and is laterally attached to the embryo axis. In Allium (Liliaceae) the cotyledon tip remains embedded in the endosperm to absorb nutrients but the cotyledon also emerges from the soil (i.e. germination is epigeal) and becomes photosynthetic. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Cotyledon. The first leaves of the embryo that serve for food digestion and food storage. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Teeth. Pointed lobes or divisions. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Leaf. Lateral organ of the stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Petal. One of the divisions on a corolla; a showy floral leaf of the perianth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Scale. Reduced leaves at the base of a shoot. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Opposite. Bearing two leaves or buds at a node on opposite sides of a stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Scutellum. Shield-shaped organ surrounding the embryo of a grass that is morphologically like a cotyledon. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)
  
  
• Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Pericarp [fruit coat]. The wall of the mature ovary or fruit. 2020 International Rules for Seed Testing (ISTA)
  
  
• Pericarp. The ovary wall. the fruit coat enclosing the seed. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Pericarp. Fruit wall. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Pericarp. The wall of a ripened ovary; the fruit coat.  Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Epicotyl. In Dicotyledons and gymnosperms, the epicotyl includes all seedling structures above the cotyledons. In species with epigeal germination (e.g. Phaseolus vulgaris), the epicotyl, cotyledons and part of the hypocotyl emerge from the soil. In species with hypogeal germination (e.g. Pisum sativum), only the epicotyl emerges, carrying the first foliage leaves above the soil surface. In these species, the epicotyl also bears one or more scale leaves. Dormant meristematic buds in the axils of these scale leaves become active if there is damage to the terminal bud. The conducting tissue of the epicotyl transfers water and nutrients from the hypocotyl and cotyledons to the leaves and terminal bud above. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Epicotyl. The stem of the embryo or young seedling above the cotyledons. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seedling. A young plant developing from the embryo of a seed. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Cotyledon. The first leaf or pair of leaves of an embryo and seedling, often a food-storage organ. The cotyledons may remain in the seed (hypogeal germination) or emerge to become the first photosynthetic organs (epigeal germination). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Cotyledons. The cotyledons are the storage structures of the embryo. They may be only a small portion of the seed in species with endosperm, perisperm or female gametophyte storage tissue, or they may occupy a large portion of the embryo when they are the primary storage tissue (e.g. Phaseolus vulgaris). In epigeal species, the cotyledons may grow quite large and become the first photosynthetic structures of the young plant. In hypogeal species the primary function of the cotyledons is to provide nutrients to the growing seedling until it can produce its own nutrients. In most species the cotyledons shrivel and drop off as their reserves are depleted. In a few species (e.g. Cucurbita pepo, pumpkin) the cotyledons may persist well beyond the seedling stage of growth. In the monocotyledons the cotyledon absorbs nutrients from the endosperm and transfers them to the growing seedling. In the Poaceae the cotyledon is called the scutellum. It is in close proximity to the endosperm and is laterally attached to the embryo axis. In Allium (Liliaceae) the cotyledon tip remains embedded in the endosperm to absorb nutrients but the cotyledon also emerges from the soil (i.e. germination is epigeal) and becomes photosynthetic. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Cotyledon. The first leaves of the embryo that serve for food digestion and food storage. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Species. A category of classification lower than a genus that is made up of plants which possess in common distinctive characteristics that are reproduced in their offspring. The species name included second to the genus in the scientific binomial. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Epigeal. Cotyledons borne above the ground after germination (see hypogeal). Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
  
  
• Epigeal germination. A type of germination in which cotyledons are carried above soil level by the elongating hypocotyl (see hypogeal germination). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Hypocotyl. The portion of the seedling axis between the root and the cotyledons is the hypocotyl. The hypocotyl is a transition structure for the transport of water and dissolved salts from the roots to the epicotyl. When a seed with epigeal germination (see section 2.5 for definitions of epigeal and hypogeal germination) is planted in moist soil, the hypocotyl elongates carrying the cotyledons above the soil surface.
  
  In monocotyledons the hypocotyl is usually not discernible as a separate structure. The mesocotyl is the part of the seedling axis between the scutellum and the base of the coleoptile. In some species (e.g. Zea mays) the elongation of the mesocotyl may be considerable. In others (e.g. Triticum aestivum) the elongation may be imperceptible. Elongation of the mesocotyl is suppressed by light after the coleoptile emerges from the soil.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Hypocotyl. That portion of the embryo stem below the cotyledons. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Hypogeal. The cotyledons borne below the ground after germination (see epigeal). Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)
  
  
• Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

• Leaf. Lateral organ of the stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Scale leaf. A reduced leaf, usually appressed to the stem (e.g. in Asparagus, Pisum). SCST Seed Technologist Training Manual, 2018

• Dormant seeds. Viable seeds, other than hard seeds, which fail to germinate when provided the specified germination conditions for the kind of seed in question. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Dormancy. Delayed germination or growth; a condition of inactivity. SCST Seed Technologist Training Manual, 2018

• Bud. An unexpanded flower or a rudimentary leaf, stem, or branch. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Scale. Reduced leaves at the base of a shoot. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Terminal bud. The shoot apex enveloped by several more or less differentiated leaves. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Terminal bud. The terminal bud occupies the tip of the epicotyl and consists of the apical meristem surrounded and protected by the developing leaves.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Conducting tissues. Tissues that transport water and dissolved minerals from the root to the other plant structures, and foods from where they are manufactured (e.g., leaves) to where they are needed for growth or storage. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Terminal. Extremity or upper bud, flower, or leaf. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Bud. An unexpanded flower or a rudimentary leaf, stem, or branch. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Embryo. The young plant enclosed in a seed. 2020 International Rules for Seed Testing (ISTA)
  
  
• Embryo. Rudimentary plant enclosed in a seed, usually consisting of a more or less differentiated axis and attached cotyledon(s). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• The embryo. The product of one of the fusions of the angiosperm fertilization process is the embryo (the other being the endosperm). In gymnosperms the embryo is the only product of the fertilization process. Depending on the species, the embryo develops to varying degrees within the seed, becoming a 'miniature plant' by the end of the growing season. In Phaseolus vulgaris, for example, the embryo is fully developed and the radicle, hypocotyl and epicotyl with primary leaves can easily be observed. The development of the embryo in other species may be much less, with some essential structures being observed only after considerable growth of the seedling. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Embryo. A young plant before the beginning if its rapid growth. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Leaf. Lateral organ of the stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Cotyledon. The first leaf or pair of leaves of an embryo and seedling, often a food-storage organ. The cotyledons may remain in the seed (hypogeal germination) or emerge to become the first photosynthetic organs (epigeal germination). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Cotyledons. The cotyledons are the storage structures of the embryo. They may be only a small portion of the seed in species with endosperm, perisperm or female gametophyte storage tissue, or they may occupy a large portion of the embryo when they are the primary storage tissue (e.g. Phaseolus vulgaris). In epigeal species, the cotyledons may grow quite large and become the first photosynthetic structures of the young plant. In hypogeal species the primary function of the cotyledons is to provide nutrients to the growing seedling until it can produce its own nutrients. In most species the cotyledons shrivel and drop off as their reserves are depleted. In a few species (e.g. Cucurbita pepo, pumpkin) the cotyledons may persist well beyond the seedling stage of growth. In the monocotyledons the cotyledon absorbs nutrients from the endosperm and transfers them to the growing seedling. In the Poaceae the cotyledon is called the scutellum. It is in close proximity to the endosperm and is laterally attached to the embryo axis. In Allium (Liliaceae) the cotyledon tip remains embedded in the endosperm to absorb nutrients but the cotyledon also emerges from the soil (i.e. germination is epigeal) and becomes photosynthetic. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Cotyledon. The first leaves of the embryo that serve for food digestion and food storage. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Germination. Germination of a seed in an ISTA test is the emergence and development of the seedling to a stage where the aspect of its essential structures indicates whether or not it is able to develop further into a satisfactory plant under favourable conditions in the field. 2020 International Rules for Seed Testing, (ISTA)
  
  
• Germination (seed testing definition).The emergence and development from the seed embryo of those essential structures which, for the kind of seed in question, are indicative of its ability to produce a normal plant under favorable conditions. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Germination (physiological definition). A process involving water uptake, metabolic changes and cell elongation resulting in radicle emergence from the seed. SCST Seed Technologist Training Manual, 2018

• Hypogeal. The cotyledons borne below the ground after germination (see epigeal). Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Hypocotyl. The portion of the seedling axis between the root and the cotyledons is the hypocotyl. The hypocotyl is a transition structure for the transport of water and dissolved salts from the roots to the epicotyl. When a seed with epigeal germination (see section 2.5 for definitions of epigeal and hypogeal germination) is planted in moist soil, the hypocotyl elongates carrying the cotyledons above the soil surface.
  
  In monocotyledons the hypocotyl is usually not discernible as a separate structure. The mesocotyl is the part of the seedling axis between the scutellum and the base of the coleoptile. In some species (e.g. Zea mays) the elongation of the mesocotyl may be considerable. In others (e.g. Triticum aestivum) the elongation may be imperceptible. Elongation of the mesocotyl is suppressed by light after the coleoptile emerges from the soil.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Hypocotyl. That portion of the embryo stem below the cotyledons. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seedling. A young plant developing from the embryo of a seed. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Normal seedling. A seedling with all essential structures present and capable of developing into a plant under favorable conditions; certain defects may be present if they are judged to be not so severe as to impede further development of the plant (see abnormal seedling). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Abnormal seedling. A seedling that does not have all the essential structures or is damaged, deformed or decayed to such an extent that normal development is prevented (see normal seedling). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Essential structure. Structure which is critical for continued development of the seedling into a plant. SCST Seed Technologist Training Manual. 2018
  
  
• Essential structure. Any seedling structure that must be considered when classifying a seedling as either normal or abnormal. In general, an essential structure is one that is critical for successful establishment and development of a seedling into a plant. Structure which is critical for continued development of the seedling into a plant. Baalbaki, R.Z. Germination & Dormancy. TWS Website. 2020

• Abnormal seedling. A seedling that does not have all the essential structures or is damaged, deformed or decayed to such an extent that normal development is prevented (see normal seedling). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Endosperm. Nutritive tissue originating from fertilization and retained at maturity in some seeds as a storage tissue for food reserves. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Endosperm.The nutritive tissue developed as a result of fertilization associated with the embryo in seeds. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
  
  
• Endosperm. Endosperm is one of the products of double fertilization, and in some species, particularly those of the Poaceae, it develops as the storage tissue. In this event, little or no nucellus tissue remains, and the extent of cotyledon development varies. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Nerve. Vascular veins of  the blades, glumes and lemmas. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Leaf. Lateral organ of the stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Branch. A lateral stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Leaf. Lateral organ of the stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Sheath. The lower part of the leaf that encloses the stem, as in grasses. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Branch. A lateral stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Cluster. A densely crowded inflorescence or, in Beta, part of an inflorescence. 2020 International Rules for Seed Testing (ISTA)

• Spikelet. The unit of a grass inflorescence comprising one or more florets subtended by one or two sterile glumes. For the purposes of the Rules, the term spikelet includes, as well as a fertile floret, either one or more additional fertile or completely infertile florets, or glumes. 2020 International Rules for Seed Testing (ISTA)
  
  
• Spikelet. Part of a grass inflorescence including one or more florets subtended by one or two glumes. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Spikelet. In grasses: One or more attached florets usually subtended by one or more bracts (glumes). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Spikelet. The unit of a grass inflorescence that consists of a pair of empty glumes that enclose one of more florets arranged around a rachilla. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Bristle. A stiff hair; sometimes applied to the upper part of an awn, when the latter is bent. 2020 International Rules for Seed Testing (ISTA)
  
  
• Bristle. A stiff hair, the upped part of an bent awn. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Bristle. A short stiff hair. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Female gametophyte. In gymnosperms the nutritive tissue is the mature female gametophyte, sometimes also referred to as the primary endosperm because it is already present before fertilization.    Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Endosperm. Nutritive tissue originating from fertilization and retained at maturity in some seeds as a storage tissue for food reserves. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Endosperm.The nutritive tissue developed as a result of fertilization associated with the embryo in seeds. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
  
  
• Endosperm. Endosperm is one of the products of double fertilization, and in some species, particularly those of the Poaceae, it develops as the storage tissue. In this event, little or no nucellus tissue remains, and the extent of cotyledon development varies. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019

• Fertilization. A sexual process in which two dissimilar gametes fuse. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Floret. General: An individual flower within a cluster. In grasses: A flower usually enclosed by two bracts (lemma and palea). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Floret. A flower within an inflorescence or in a grass spikelet. SCST Seed Technologist Training Manual, 2018
  
  
• Floret. The lemma and palea with enclosed pistil and stamens, or, in Poaceae, the mature caryopsis; for the purpose of the Rules, the term floret refers to the fertile floret, with or without additional sterile lemmas. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Floret. The lemma and palea with enclosed pistil and stamens or the mature caryopsis in Poaceae; for the purpose of the Rules, the term floret refers to the fertile floret with or without additional sterile lemmas. 2020 International Rules for Seed Testing (ISTA)
  
  
• Floret. In grasses, a flower consisting of lemma and palea which enclose the flowering parts -- stamens, pistil and lodicules. May be perfect, staminate, pistillate, neuter, sterile, and so on. A small flower in other plant families. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Caryopsis. In grasses: A dry, one-seeded, indehiscent fruit with the pericarp fused to the seed coat (testa). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Caryopsis. Naked grass-fruit in which the testa is united with the pericarp. 2020 International Rules for Seed Testing (ISTA)
  
  
• Caryopsis. A naked grass fruit in which the testa is fused with the pericarp; also a grain. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Caryopsis. The fruit of a grass, which is dry, one-seeded, indehiscent, with the testa and the pericarp completely united. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fruit. The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Fruit. In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity. SCST Seed Technologist Training Manual, 2018
  
  
• Fruit. The ripened ovary of a seed plant and associated parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fertile floret. A floret that encloses a caryopsis. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

• Pistillate. Having a pistil. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Floret. General: An individual flower within a cluster. In grasses: A flower usually enclosed by two bracts (lemma and palea). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Floret. A flower within an inflorescence or in a grass spikelet. SCST Seed Technologist Training Manual, 2018
  
  
• Floret. The lemma and palea with enclosed pistil and stamens, or, in Poaceae, the mature caryopsis; for the purpose of the Rules, the term floret refers to the fertile floret, with or without additional sterile lemmas. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Floret. The lemma and palea with enclosed pistil and stamens or the mature caryopsis in Poaceae; for the purpose of the Rules, the term floret refers to the fertile floret with or without additional sterile lemmas. 2020 International Rules for Seed Testing (ISTA)
  
  
• Floret. In grasses, a flower consisting of lemma and palea which enclose the flowering parts -- stamens, pistil and lodicules. May be perfect, staminate, pistillate, neuter, sterile, and so on. A small flower in other plant families. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Caryopsis. In grasses: A dry, one-seeded, indehiscent fruit with the pericarp fused to the seed coat (testa). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Caryopsis. Naked grass-fruit in which the testa is united with the pericarp. 2020 International Rules for Seed Testing (ISTA)
  
  
• Caryopsis. A naked grass fruit in which the testa is fused with the pericarp; also a grain. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Caryopsis. The fruit of a grass, which is dry, one-seeded, indehiscent, with the testa and the pericarp completely united. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Root. In dicotyledons and gymnosperms, the root system serves three major functions: (1) to anchor the plant in the soil, (2) to absorb water and dissolved salts from the soil and (3) to conduct the water and salts to the hypocotyl, cotyledons and epicotyl. The embryonic root, or radicle, is located at the basal end of the embryo and is usually the first seedling structure to rupture the testa. After emergence it is referred to as the primary root. The primary root elongates rapidly and soon numerous root hairs develop, greatly increasing the absorbing surface of the roots. As the seedling continues to grow, secondary roots develop from the primary root and from other secondary roots. Roots may also emerge from other structures (e.g. the hypocotyl) and are referred to as adventitious roots. As in the dicotyledons, the monocotyledon root system serves to anchor the plant in soil, absorb water and dissolved salts from the soil and to conduct the water and salts to the growing seedling. The embryonic root, or radicle, is situated at the basal end of the embryo and, in the case of the Poaceae, its apex is covered by the coleorhiza. After the radicle emerges it is referred to as the primary root. In some species of the Poaceae (e.g. Triticum) the primary root is indistinguishable from the other roots that develop from the scutellar node region and hence all of these are referred to as seminal roots. Roots that develop from structures above the scutellar or cotyledonary node are called adventitious roots. Secondary roots may develop from seminal and adventitious roots.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Root. A portion of a higher plant bearing neither leaves nor reproductive organs, usually underground. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Root. In dicotyledons and gymnosperms, the root system serves three major functions: (1) to anchor the plant in the soil, (2) to absorb water and dissolved salts from the soil and (3) to conduct the water and salts to the hypocotyl, cotyledons and epicotyl. The embryonic root, or radicle, is located at the basal end of the embryo and is usually the first seedling structure to rupture the testa. After emergence it is referred to as the primary root. The primary root elongates rapidly and soon numerous root hairs develop, greatly increasing the absorbing surface of the roots. As the seedling continues to grow, secondary roots develop from the primary root and from other secondary roots. Roots may also emerge from other structures (e.g. the hypocotyl) and are referred to as adventitious roots. As in the dicotyledons, the monocotyledon root system serves to anchor the plant in soil, absorb water and dissolved salts from the soil and to conduct the water and salts to the growing seedling. The embryonic root, or radicle, is situated at the basal end of the embryo and, in the case of the Poaceae, its apex is covered by the coleorhiza. After the radicle emerges it is referred to as the primary root. In some species of the Poaceae (e.g. Triticum) the primary root is indistinguishable from the other roots that develop from the scutellar node region and hence all of these are referred to as seminal roots. Roots that develop from structures above the scutellar or cotyledonary node are called adventitious roots. Secondary roots may develop from seminal and adventitious roots.  Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Root. A portion of a higher plant bearing neither leaves nor reproductive organs, usually underground. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Stalk. The stem of any plant organ. 2020 International Rules for Seed Testing (ISTA)

• Anther. The pollen-producing part of the stamen, borne at the top of the filament or stalk. 2020 International Rules for Seed Testing, (ISTA)
  
  
• Anther. Pollen-bearing part of stamen. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Stamen. The male sex organ of a flower, including the filament and anther. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Stamen.The pollen-bearing organs of a flower. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Opposite. Bearing two leaves or buds at a node on opposite sides of a stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Turgid. Swollen. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
  
  
• Turgid cells. Each healthy turgid cell is swollen due to high turgor pressure. Tissues made of of turgid cells that stain normally are considered healthy and functional. TWS
  
  
•  Turgid. A balloon that is full of air is turgid. TWS

Flaccid. Cells in plant or seed tissue that are soft. The opposite of turgid, or swollen.  TWS

Flaccid. A ballon that is losing air is becoming flaccid. TWS

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fleshy fruits. Fruits that retain moisture. Seeds within fleshy fruits remain at a high moisture content throughout seed development. Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Opposite. Bearing two leaves or buds at a node on opposite sides of a stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Cluster. A densely crowded inflorescence or, in Beta, part of an inflorescence. 2020 International Rules for Seed Testing (ISTA)

• Lemma. The outer (lower) bract of a grass floret, sometimes referred to as the flowering glume or the lower or outer palea. Bract enclosing the caryopsis on the outer (dorsal) side. 2020 International Rules for Seed Testing (ISTA)
  
  
• Lemma. The outer (lower) bract of a grass floret; enclosing the caryopsis on the outer (dorcal) side. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Lemma. The lower of two bracts that subtend a grass flower. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Lemma. The lower of the two bracts enclosing the flower in the spikelet of grasses. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Palea. The upper (inner) bract of a grass floret, sometimes called the inner or upper palea. Bract enclosing the caryopsis on the inner ventral side. 2020 International Rules for Seed Testing (ISTA)
  
  
• Palea. The upper (inner) bract of a grass floret, the bract enclosing the caryopsis on the inner (ventral) side. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Palea. The uppermost of two bracts subtending a grass flower. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Palea. The upper of the two bracts enclosing the flower in the spikelet of grasses.  Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Inflorescence. A flower cluster. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Inflorescence. The portion of the plant adapted for flowering and fruiting. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Spikelet. The unit of a grass inflorescence comprising one or more florets subtended by one or two sterile glumes. For the purposes of the Rules, the term spikelet includes, as well as a fertile floret, either one or more additional fertile or completely infertile florets, or glumes. 2020 International Rules for Seed Testing (ISTA)
  
  
• Spikelet. Part of a grass inflorescence including one or more florets subtended by one or two glumes. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Spikelet. In grasses: One or more attached florets usually subtended by one or more bracts (glumes). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Spikelet. The unit of a grass inflorescence that consists of a pair of empty glumes that enclose one of more florets arranged around a rachilla. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Pistil.  A part (or whole) of the gynoecium, consisting of either a separate, free carpel or two or more fused carpels. A typical pistil comprises an ovary, a style and a stigma. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Pistil. The ovule-bearing portion of a flower consisting of a stigma, style and ovary. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Caryopsis. In grasses: A dry, one-seeded, indehiscent fruit with the pericarp fused to the seed coat (testa). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Caryopsis. Naked grass-fruit in which the testa is united with the pericarp. 2020 International Rules for Seed Testing (ISTA)
  
  
• Caryopsis. A naked grass fruit in which the testa is fused with the pericarp; also a grain. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Caryopsis. The fruit of a grass, which is dry, one-seeded, indehiscent, with the testa and the pericarp completely united. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Floret. General: An individual flower within a cluster. In grasses: A flower usually enclosed by two bracts (lemma and palea). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Floret. A flower within an inflorescence or in a grass spikelet. SCST Seed Technologist Training Manual, 2018
  
  
• Floret. The lemma and palea with enclosed pistil and stamens, or, in Poaceae, the mature caryopsis; for the purpose of the Rules, the term floret refers to the fertile floret, with or without additional sterile lemmas. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Floret. The lemma and palea with enclosed pistil and stamens or the mature caryopsis in Poaceae; for the purpose of the Rules, the term floret refers to the fertile floret with or without additional sterile lemmas. 2020 International Rules for Seed Testing (ISTA)
  
  
• Floret. In grasses, a flower consisting of lemma and palea which enclose the flowering parts -- stamens, pistil and lodicules. May be perfect, staminate, pistillate, neuter, sterile, and so on. A small flower in other plant families. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fertile floret. A floret that encloses a caryopsis. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

• Sterile. Without functional sex organs (for grass florets: without caryopsis). 2020 International Rules for Seed Testing (ISTA)
  
  
• Sterile. Without pistils. A sterile floret may be staminate or neuter. It may even lack a palea, and consist of nothing but a lemma. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Lemma. The outer (lower) bract of a grass floret, sometimes referred to as the flowering glume or the lower or outer palea. Bract enclosing the caryopsis on the outer (dorsal) side. 2020 International Rules for Seed Testing (ISTA)
  
  
• Lemma. The outer (lower) bract of a grass floret; enclosing the caryopsis on the outer (dorcal) side. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Lemma. The lower of two bracts that subtend a grass flower. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Lemma. The lower of the two bracts enclosing the flower in the spikelet of grasses. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fertile. With functional sex organs; (for grass florets: having a caryopsis). 2020 International Rules for Seed Testing (ISTA)
  
  
• Fertile. With functional sex organs. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Fertile. Capable of producing fruit; having pistils. A fertile floret may be pistillate or perfect. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Lodicule. The organs at the base of the ovary of a grass flower that swell and open the lemma and palea during anthesis. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111
  
  
• Lodicules. Scale-like structures in a grass flower that swell and force open the surrounding structures to facilitate pollination. SCST Seed Technologist Training Manual, 2018

• Staminate. Flower with stamens only. 2020 International Rules for Seed Testing (ISTA)
  
  
• Staminate. Containing stamens only. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Pistillate. Having a pistil. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Neuter. Without stamens or pistils. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Dense. Parts massed or crowded together. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Cluster. A densely crowded inflorescence or, in Beta, part of an inflorescence. 2020 International Rules for Seed Testing (ISTA)

• Sessile. Without a stalk or pedicel. 2020 International Rules for Seed Testing (ISTA)
  
  
• Sessile. Attached directly by the base and not raised on a stalk. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Axis. The central line of any organ or the support of a group of organs; the main stem of an inflorescence, especially of a panicle. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Capitulum, capitula (plural). A dense inflorescence of usually sessile flowers. 2020 International Rules for Seed Testing (ISTA)
  
  
• Capitulum, capitula (plural). A compact inforescence with a disc of sessile flowers, e.g. Asteraceae (flower head). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010.
  
  
• Capitulum, capitula (plural). A small head inflorescence. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019

• Dehiscent fruit. A dry fruit, that experiences desiccation. Seeds within dehiscent fruits will lose moisture content during the later stages of seed development. Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020
  
  
• Dehiscent fruit. A fruit that opens at maturity allowing seeds to be released from the fruit. SCST Seed Technologist Training Manual, 2018
  
  
• Dehiscent. A fruit in which the fruit wall splits open at maturity to release the seed. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Dehiscent. Opening spontaneously at maturity. SCST Seed Technologist Training Manual, 2018

• Fruit. The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Fruit. In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity. SCST Seed Technologist Training Manual, 2018
  
  
• Fruit. The ripened ovary of a seed plant and associated parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Carpel. The female reproductive organ of flowering plants. It consists of an ovary, containing one or more ovules (which became seeds after fertilization), and a stigma, a surface receptive for pollen grains. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Carpel. A simple pistil or an element of a compound pistil. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Ventral. The side facing towards the axis; the lower surface (dorsal). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Ventral. The lower or front side; opposite to dorsal. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Suture. A seam or line of union. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Pericarp [fruit coat]. The wall of the mature ovary or fruit. 2020 International Rules for Seed Testing (ISTA)
  
  
• Pericarp. The ovary wall. the fruit coat enclosing the seed. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Pericarp. Fruit wall. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Pericarp. The wall of a ripened ovary; the fruit coat.  Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Succulent. Fruits in which the mesocarp develops into a fleshy covering. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Succulent. Fleshy or juicy. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Mesocarp. The middle layer of the pericarp. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Mesocarp. The middle layer of the pericarp (fruit wall) between the endocarp and exocarp. SCST Seed Technologist Training Manual, 2018

• Dehiscent fruit. A dry fruit, that experiences desiccation. Seeds within dehiscent fruits will lose moisture content during the later stages of seed development. Bowden, L., Moisture Testing in Wild Species. TWS Website. 2020
  
  
• Dehiscent fruit. A fruit that opens at maturity allowing seeds to be released from the fruit. SCST Seed Technologist Training Manual, 2018
  
  
• Dehiscent. A fruit in which the fruit wall splits open at maturity to release the seed. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Dehiscent. Opening spontaneously at maturity. SCST Seed Technologist Training Manual, 2018

• Indehiscent. Not opening; fruits which do not open at maturity. 2020 International Rules for Seed Testing (ISTA)
  
  
• Indehiscent. Fruits in which the fruit wall does not split open at maturity to release the seed (see dehiscent). ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Indehiscent fruit. A fruit that does not open at maturity. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Indehiscent. Fruits remaining closed at maturity. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fruit. The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Fruit. In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity. SCST Seed Technologist Training Manual, 2018
  
  
• Fruit. The ripened ovary of a seed plant and associated parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Open. Loose; opposite of dense or compact. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Gynoecium. The female reproductive part of a flower, composed of one or more pistils. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Gynoecium. The carpels taken collectively. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Inflorescence. A flower cluster. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Inflorescence. The portion of the plant adapted for flowering and fruiting. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Monocarpellary. Derived from a single ovary. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

• Polycarpellary. Seeds derived from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

• Ovary. The basal, expanded part of the pistil that contains the ovules. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Ovary. The basal enlarged portion of a pistil within which seeds develop. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Angiosperm. A plant whose seeds are borne within a mature ovary (fruit). SCST Seed Technologist Training Manual, 2018

• Rudimentary. Imperfectly developed. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Leaf. Lateral organ of the stem. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Fruit. The structure that develops from the pericarp as the enclosed seed or seeds mature. Fruits may have the following attributes: Succulent or dry, depending on whether or not the middle layer of the pericarp (mesocarp) develops into a fleshy covering; Dehiscent or indehiscent, according to whether or not the fruit wall splits open to release the seed; True or simple fruits, which develop from the gynoecium of a single flower, and multiple fruits, which develop from a complete inflorescence; Monocarpellary or polycarpellary, depending on whether they developed from a single ovary or from a number of fused ovaries. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Fruit. In angiosperms, a mature ripened ovary, usually containing seeds. Some authors include extracarpellary parts adhering to the ovary at maturity. SCST Seed Technologist Training Manual, 2018
  
  
• Fruit. The ripened ovary of a seed plant and associated parts. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Funiculus. The stalk of an ovule. Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 1, 2019
  
  
• Funicle, funiculus. The cord or thread which connects the ovule and later the seed to the placenta. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Stalk. The stem of any plant organ. 2020 International Rules for Seed Testing (ISTA)

• Ovule. The immature seed within the ovary. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Ovule. A structure, consisting of a female gametophyte, nucellus, and integuments, which develops into a seed after fertilization. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Seed. The part of a plant which is able to develop into a new plant. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010
  
  
• Seed. Botanically, a seed is a mature fertilized ovule containing an embryonic plant; usually it has nutrient storage tissue and is surrounded by a protective coat, the testa. This structure is a "true seed"; however, the ovules of many species have additional structures of the mother plant attached or fused to the seed coat. For example, the "seed" of Triticum aestivum (wheat) is botanically a fruit because the pericarp (ovary wall) is fused with the seed coat. In these rules the term "seed" will be used in the agronomic sense (i.e. the true seed plus any accessory structures that may be attached when it is planted in the field; see section 3.2 of the AOSA Rules for Testing Seeds Vol. 1). Association of Official Seed Analysts (AOSA) Rules for Seed Testing, Volume 4, 2019
  
  
• Seed. The ripened ovule, enclosing the rudimentary plant and food necessary for its germination. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Placenta. The part of the ovary where the funicles attach. ISTA Handbook on Pure Seed Definitions, Glossary. 3rd Edition 2010

• Gamete. A matured sex cell capable of uniting with another gamete to form a new individual. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Species. A category of classification lower than a genus that is made up of plants which possess in common distinctive characteristics that are reproduced in their offspring. The species name included second to the genus in the scientific binomial. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111

• Genus. A group of closely related species. The genus name is the first word of a binomial scientific name and is capitalized. The plural form of the word is genera. Fenwick, J.R. revised 1995. Laboratory Manual for General Crops- Glossary. Unpublished class notes. Dept. of Soil and Crop Sciences, Colorado State University. Pages 104-111